{"id":"https://openalex.org/W7225617762","doi":"https://doi.org/10.3389/fpls.2026.1986895","title":"Tomato cultivation substrates differ in stage-dependent root zone\u2013rhizosphere coordination and biomass allocation","display_name":"Tomato cultivation substrates differ in stage-dependent root zone\u2013rhizosphere coordination and biomass allocation","publication_year":2026,"publication_date":"2026-11-09","ids":{"openalex":"https://openalex.org/W7225617762","doi":"https://doi.org/10.3389/fpls.2026.1986895"},"language":"en","primary_location":{"id":"doi:10.3389/fpls.2026.1986895","doi":"https://doi.org/10.3389/fpls.2026.1986895","is_oa":true,"landing_page_url":"https://doi.org/10.3389/fpls.2026.1986895","pdf_url":"https://www.frontiersin.org/journals/plant-science/articles/10.3389/fpls.2026.1986895/pdf","source":{"id":"https://openalex.org/S2493613807","display_name":"Frontiers in Plant Science","issn_l":"1664-462X","issn":["1664-462X"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","norway-2","poland-100","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310320527","host_organization_name":"Frontiers Media","host_organization_lineage":["https://openalex.org/P4310320527"],"host_organization_lineage_names":["Frontiers Media"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Frontiers in Plant Science","raw_type":"journal-article"},"volume":null,"type":"article","is_traditional":true,"indexed_in":["crossref","doaj"],"open_access":{"is_oa":true,"oa_status":"gold","oa_url":"https://www.frontiersin.org/journals/plant-science/articles/10.3389/fpls.2026.1986895/pdf","any_repository_has_fulltext":false},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5100687984","display_name":"Yu Chen","orcid":"https://orcid.org/0000-0002-7258-2607","observed_orcids":["https://orcid.org/0000-0002-7258-2607"]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Yu Chen","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University","Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]},{"raw_affiliation_string":"Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5130939789","display_name":"Yunxue Li","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Yunxue Li","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University","Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]},{"raw_affiliation_string":"Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5130975549","display_name":"Wenyin Zi","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Wenyin Zi","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University","Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]},{"raw_affiliation_string":"Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5132622451","display_name":"Yuyuan Chai","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Yuyuan Chai","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5158664365","display_name":"Jianjun Chen","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":true,"raw_author_name":"Jianjun Chen","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]},{"author_position":"last","author":{"id":"https://openalex.org/A5161326202","display_name":"Jing Shi","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I195019228","display_name":"Yunnan Agricultural University","ror":"https://ror.org/04dpa3g90","country_code":"CN","type":"education","lineage":["https://openalex.org/I195019228"]}],"countries":["CN"],"is_corresponding":true,"raw_author_name":"Jing Shi","raw_affiliation_strings":["College of Resources and Environment, Yunnan Agricultural University","Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]},{"raw_affiliation_string":"Key Laboratory for Improving Quality and Productivity of Arable Land of Yunnan Province, College of Resources and Environment, Yunnan Agricultural University","institution_ids":["https://openalex.org/I195019228"]}]}],"institutions":[],"countries_distinct_count":1,"institutions_distinct_count":1,"corresponding_author_ids":["https://openalex.org/A5158664365","https://openalex.org/A5161326202"],"corresponding_institution_ids":["https://openalex.org/I195019228"],"apc_list":{"value":3150,"currency":"CHF","value_usd":3801},"apc_paid":{"value":3150,"currency":"CHF","value_usd":3801},"fwci":0.0,"has_fulltext":true,"cited_by_count":0,"citation_normalized_percentile":{"value":0.0,"is_in_top_1_percent":false,"is_in_top_10_percent":false},"cited_by_percentile_year":null,"biblio":{"volume":"17","issue":null,"first_page":null,"last_page":null},"is_retracted":false,"is_paratext":false,"is_xpac":false,"primary_topic":{"id":"https://openalex.org/T12113","display_name":"Growth and nutrition in plants","score":0.9384999871253967,"subfield":{"id":"https://openalex.org/subfields/1110","display_name":"Plant Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},"topics":[{"id":"https://openalex.org/T12113","display_name":"Growth and nutrition in plants","score":0.9384999871253967,"subfield":{"id":"https://openalex.org/subfields/1110","display_name":"Plant Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T12754","display_name":"Plant Growth and Agriculture Techniques","score":0.030700000002980232,"subfield":{"id":"https://openalex.org/subfields/1111","display_name":"Soil Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T12605","display_name":"Berry genetics and cultivation research","score":0.009600000455975533,"subfield":{"id":"https://openalex.org/subfields/1110","display_name":"Plant Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[{"id":"https://openalex.org/keywords/rhizosphere","display_name":"rhizosphere","score":0.6330000162124634},{"id":"https://openalex.org/keywords/biomass-allocation","display_name":"biomass allocation","score":0.7039999961853027},{"id":"https://openalex.org/keywords/developmental-stages","display_name":"developmental stages","score":0.6169999837875366},{"id":"https://openalex.org/keywords/peat-substrate","display_name":"peat substrate","score":0.9760000109672546},{"id":"https://openalex.org/keywords/rhizosphere-enzyme-activity","display_name":"rhizosphere enzyme activity","score":0.8679999709129333},{"id":"https://openalex.org/keywords/dehydrogenase-activity","display_name":"dehydrogenase activity","score":0.9440000057220459},{"id":"https://openalex.org/keywords/urease","display_name":"urease","score":0.7680000066757202},{"id":"https://openalex.org/keywords/sucrase","display_name":"sucrase","score":0.8870000243186951},{"id":"https://openalex.org/keywords/fluorescein-diacetate-hydrolase","display_name":"fluorescein diacetate hydrolase","score":0.9599999785423279},{"id":"https://openalex.org/keywords/root-traits","display_name":"root traits","score":0.6880000233650208},{"id":"https://openalex.org/keywords/nutrient-accumulation","display_name":"nutrient accumulation","score":0.5450000166893005},{"id":"https://openalex.org/keywords/yield","display_name":"yield","score":0.5419999957084656},{"id":"https://openalex.org/keywords/protected-cultivation","display_name":"protected cultivation","score":0.4090000092983246}],"concepts":[{"id":"https://openalex.org/C86626682","wikidata":"https://www.wikidata.org/wiki/Q1124578","display_name":"Rhizosphere","level":3,"score":0.8860999941825867},{"id":"https://openalex.org/C2780138947","wikidata":"https://www.wikidata.org/wiki/Q393855","display_name":"Dry matter","level":2,"score":0.5953999757766724},{"id":"https://openalex.org/C6557445","wikidata":"https://www.wikidata.org/wiki/Q173113","display_name":"Agronomy","level":1,"score":0.5820000171661377},{"id":"https://openalex.org/C185592680","wikidata":"https://www.wikidata.org/wiki/Q2329","display_name":"Chemistry","level":0,"score":0.5266000032424927},{"id":"https://openalex.org/C2781083041","wikidata":"https://www.wikidata.org/wiki/Q146555","display_name":"Solanum","level":2,"score":0.519599974155426},{"id":"https://openalex.org/C142796444","wikidata":"https://www.wikidata.org/wiki/Q181394","display_name":"Nutrient","level":2,"score":0.4893999993801117},{"id":"https://openalex.org/C115540264","wikidata":"https://www.wikidata.org/wiki/Q2945560","display_name":"Biomass (ecology)","level":2,"score":0.43720000982284546},{"id":"https://openalex.org/C2777289219","wikidata":"https://www.wikidata.org/wiki/Q7632154","display_name":"Substrate (aquarium)","level":2,"score":0.4327000081539154},{"id":"https://openalex.org/C144027150","wikidata":"https://www.wikidata.org/wiki/Q48803","display_name":"Horticulture","level":1,"score":0.41909998655319214},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.3763999938964844},{"id":"https://openalex.org/C53657456","wikidata":"https://www.wikidata.org/wiki/Q184624","display_name":"Peat","level":2,"score":0.36959999799728394},{"id":"https://openalex.org/C150668497","wikidata":"https://www.wikidata.org/wiki/Q104870826","display_name":"Dry weight","level":2,"score":0.3546999990940094},{"id":"https://openalex.org/C48743137","wikidata":"https://www.wikidata.org/wiki/Q1783121","display_name":"Organic matter","level":2,"score":0.3513999879360199},{"id":"https://openalex.org/C59822182","wikidata":"https://www.wikidata.org/wiki/Q441","display_name":"Botany","level":1,"score":0.33379998803138733},{"id":"https://openalex.org/C2776317432","wikidata":"https://www.wikidata.org/wiki/Q412547","display_name":"Dehydrogenase","level":3,"score":0.3068999946117401},{"id":"https://openalex.org/C137580998","wikidata":"https://www.wikidata.org/wiki/Q235352","display_name":"Crop","level":2,"score":0.30390000343322754},{"id":"https://openalex.org/C183889291","wikidata":"https://www.wikidata.org/wiki/Q5187955","display_name":"Crop residue","level":3,"score":0.2883000075817108},{"id":"https://openalex.org/C159750122","wikidata":"https://www.wikidata.org/wiki/Q96621023","display_name":"Soil water","level":2,"score":0.2881999909877777},{"id":"https://openalex.org/C2781174152","wikidata":"https://www.wikidata.org/wiki/Q65772506","display_name":"Sinigrin","level":4,"score":0.2689000070095062},{"id":"https://openalex.org/C157517311","wikidata":"https://www.wikidata.org/wiki/Q167980","display_name":"Transpiration","level":3,"score":0.2637999951839447},{"id":"https://openalex.org/C137660486","wikidata":"https://www.wikidata.org/wiki/Q732240","display_name":"Growing season","level":2,"score":0.2624000012874603},{"id":"https://openalex.org/C28212737","wikidata":"https://www.wikidata.org/wiki/Q5381543","display_name":"Enzyme assay","level":3,"score":0.2612000107765198},{"id":"https://openalex.org/C2780777981","wikidata":"https://www.wikidata.org/wiki/Q3266947","display_name":"Beta-glucosidase","level":4,"score":0.2506999969482422}],"mesh":[],"locations_count":1,"locations":[{"id":"doi:10.3389/fpls.2026.1986895","doi":"https://doi.org/10.3389/fpls.2026.1986895","is_oa":true,"landing_page_url":"https://doi.org/10.3389/fpls.2026.1986895","pdf_url":"https://www.frontiersin.org/journals/plant-science/articles/10.3389/fpls.2026.1986895/pdf","source":{"id":"https://openalex.org/S2493613807","display_name":"Frontiers in Plant Science","issn_l":"1664-462X","issn":["1664-462X"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","norway-2","poland-100","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310320527","host_organization_name":"Frontiers Media","host_organization_lineage":["https://openalex.org/P4310320527"],"host_organization_lineage_names":["Frontiers Media"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Frontiers in Plant Science","raw_type":"journal-article"}],"best_oa_location":{"id":"doi:10.3389/fpls.2026.1986895","doi":"https://doi.org/10.3389/fpls.2026.1986895","is_oa":true,"landing_page_url":"https://doi.org/10.3389/fpls.2026.1986895","pdf_url":"https://www.frontiersin.org/journals/plant-science/articles/10.3389/fpls.2026.1986895/pdf","source":{"id":"https://openalex.org/S2493613807","display_name":"Frontiers in Plant Science","issn_l":"1664-462X","issn":["1664-462X"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","norway-2","poland-100","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310320527","host_organization_name":"Frontiers Media","host_organization_lineage":["https://openalex.org/P4310320527"],"host_organization_lineage_names":["Frontiers Media"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Frontiers in Plant Science","raw_type":"journal-article"},"sustainable_development_goals":[{"display_name":"Zero hunger","score":0.991100013256073,"id":"https://metadata.un.org/sdg/2"}],"study_designs":[],"awards":[],"funders":[{"id":"https://openalex.org/F4320325440","display_name":"Yunnan Provincial Science and Technology Department","ror":"https://ror.org/03whqdf70"}],"has_content":{"pdf":true,"grobid_xml":true},"content_urls":{"pdf":"https://content.openalex.org/works/W7225617762.pdf","grobid_xml":"https://content.openalex.org/works/W7225617762.grobid-xml"},"referenced_works_count":69,"referenced_works":["https://openalex.org/W1598319948","https://openalex.org/W1966066251","https://openalex.org/W1971981494","https://openalex.org/W2062013915","https://openalex.org/W2074348475","https://openalex.org/W2078719939","https://openalex.org/W2315559153","https://openalex.org/W2415674567","https://openalex.org/W2741449026","https://openalex.org/W2751954542","https://openalex.org/W2793476021","https://openalex.org/W2934343392","https://openalex.org/W2969891391","https://openalex.org/W3093416821","https://openalex.org/W3096553354","https://openalex.org/W3098811216","https://openalex.org/W3122219831","https://openalex.org/W3175279414","https://openalex.org/W3200657627","https://openalex.org/W3203377449","https://openalex.org/W4200053742","https://openalex.org/W4206549489","https://openalex.org/W4211211813","https://openalex.org/W4221099747","https://openalex.org/W4224820443","https://openalex.org/W4281680997","https://openalex.org/W4283379777","https://openalex.org/W4309476506","https://openalex.org/W4310350226","https://openalex.org/W4313409431","https://openalex.org/W4313472737","https://openalex.org/W4313571281","https://openalex.org/W4318333054","https://openalex.org/W4322724357","https://openalex.org/W4323535823","https://openalex.org/W4362469401","https://openalex.org/W4376958557","https://openalex.org/W4384204152","https://openalex.org/W4385516466","https://openalex.org/W4386086101","https://openalex.org/W4386566616","https://openalex.org/W4387047728","https://openalex.org/W4388015424","https://openalex.org/W4389726908","https://openalex.org/W4389778803","https://openalex.org/W4390534926","https://openalex.org/W4390790247","https://openalex.org/W4391731458","https://openalex.org/W4391874423","https://openalex.org/W4392124947","https://openalex.org/W4392510058","https://openalex.org/W4393032696","https://openalex.org/W4393231366","https://openalex.org/W4395038505","https://openalex.org/W4396862054","https://openalex.org/W4401114875","https://openalex.org/W4401505841","https://openalex.org/W4403428976","https://openalex.org/W4406272519","https://openalex.org/W4406682086","https://openalex.org/W4407164263","https://openalex.org/W4411857434","https://openalex.org/W4412424356","https://openalex.org/W4414862893","https://openalex.org/W4415276595","https://openalex.org/W7126226017","https://openalex.org/W7147603045","https://openalex.org/W7163393032","https://openalex.org/W7171500454"],"related_works":["https://openalex.org/W2386636523","https://openalex.org/W2360914050","https://openalex.org/W3033200928","https://openalex.org/W177529285","https://openalex.org/W2390488186","https://openalex.org/W4320864235","https://openalex.org/W2391877535","https://openalex.org/W2348658073","https://openalex.org/W6920747631","https://openalex.org/W2375413563"],"abstract_inverted_index":{"Introduction":[0],"Cultivation":[1],"substrates":[2,32],"are":[3],"commonly":[4],"evaluated":[5],"from":[6],"their":[7],"initial":[8],"properties":[9,311],"or":[10,63],"final":[11],"crop":[12,304],"performance,":[13],"leaving":[14],"the":[15,165,182,293],"developmental":[16,251,317],"relationships":[17],"among":[18,296],"root-zone":[19,259,278,297],"conditions,":[20,90,298],"rhizosphere":[21,91,104,184,299],"functioning,":[22,301],"and":[23,82,88,96,113,116,118,136,153,162,174,189,195,212,238,250,261,264,302,318,327,338,340],"plant":[24,197],"responses":[25],"poorly":[26],"resolved.":[27],"We":[28],"examined":[29],"how":[30],"contrasting":[31],"were":[33,100,121,128,255],"associated":[34],"with":[35,68,204,274],"changes":[36],"in":[37,53,148,287,343],"these":[38],"components":[39],"across":[40,102,324],"tomato":[41,325,345],"development.":[42,141],"Methods":[43],"Tomato":[44],"(":[45],"Solanum":[46],"lycopersicum":[47],"L.":[48],"cv.":[49],"\u201c22B\u201d)":[50],"was":[51,75,146],"grown":[52],"greenhouse":[54],"soil":[55],"(SL),":[56],"a":[57,64,284,316,329],"composted":[58],"pinecone":[59],"residue":[60],"substrate":[61,66,249,310,336],"(PR),":[62],"peat":[65],"(PS),":[67],"corresponding":[69],"unplanted":[70],"controls.":[71],"Independent":[72],"destructive":[73],"sampling":[74],"conducted":[76],"at":[77,123,150,159,187,215],"Weeks":[78,151,160],"4,":[79],"8,":[80,154],"12,":[81],"16":[83],"after":[84],"transplanting.":[85],"Root-zone":[86],"physicochemical":[87],"nutrient":[89,341],"enzyme":[92],"activities,":[93],"root":[94,265],"traits,":[95],"organ":[97],"dry":[98,198],"matter":[99,199],"assessed":[101],"development;":[103],"dehydrogenase":[105,185],"activity,":[106],"aboveground":[107,206],"NO":[108,207],"3":[109,208],"--":[110,209],"N,":[111,210],"P,":[112,211],"K":[114,213],"accumulation":[115,214],"allocation,":[117],"yield-related":[119],"traits":[120],"determined":[122],"maturity.":[124,216],"Results":[125],"Substrate":[126],"effects":[127],"strongly":[129],"stage-dependent.":[130],"PR":[131,149,241],"generally":[132,169],"exhibited":[133,181],"higher":[134,175],"urease":[135],"sucrase":[137],"activities":[138],"during":[139,200],"early-to-middle":[140],"Fluorescein":[142],"diacetate":[143],"hydrolase":[144],"activity":[145,186],"highest":[147,183],"4":[152],"but":[155],"shifted":[156],"to":[157],"PS":[158,168,221],"12":[161],"16.":[163],"Throughout":[164],"growth":[166,326],"cycle,":[167],"maintained":[170],"lower":[171],"bulk":[172],"density":[173],"gravimetric":[176],"water":[177,339],"content.":[178],"It":[179],"also":[180],"maturity":[188],"accumulated":[190],"more":[191],"stem,":[192],"leaf,":[193],"fruit,":[194],"total":[196,268],"late":[201],"development,":[202],"together":[203],"greater":[205],"The":[217],"estimated":[218],"yield":[219],"under":[220],"reached":[222],"53,657.70":[223],"kg":[224],"ha":[225],"-1":[226,236,244],",":[227],"representing":[228],"increases":[229],"of":[230,321,335],"80.08%":[231],"over":[232,240],"SL":[233],"(29,796.45":[234],"kg\u00b7ha":[235,243],")":[237],"39.64%":[239],"(38,425.10":[242],").":[245],"After":[246],"accounting":[247],"for":[248,332],"stage,":[252],"significant":[253,272],"associations":[254],"concentrated":[256],"primarily":[257],"between":[258],"conditions":[260],"belowground":[262],"biochemical":[263,300],"responses,":[266],"whereas":[267],"biomass":[269],"showed":[270],"no":[271],"association":[273],"any":[275],"single":[276],"measured":[277],"variable.":[279],"Discussion":[280],"These":[281],"findings":[282],"support":[283],"developmental-coupling":[285],"perspective":[286,314],"which":[288],"cultivation-substrate":[289],"performance":[290],"depends":[291],"on":[292,308],"temporal":[294],"coordination":[295],"stage-specific":[303,333],"demand":[305],"rather":[306],"than":[307],"static":[309],"alone.":[312],"This":[313],"provides":[315],"process-level":[319],"understanding":[320],"substrate-rhizosphere-plant":[322],"interactions":[323],"offers":[328],"conceptual":[330],"basis":[331],"optimization":[334],"selection":[337],"management":[342],"protected":[344],"production.":[346]},"counts_by_year":[],"updated_date":"2026-11-11T12:16:26.678958","created_date":"2026-11-10T00:00:00"}