{"id":"https://openalex.org/W7214123724","doi":"https://doi.org/10.64898/2026.09.21.753065","title":"Brassinosteroid receptors reveal conserved features of exon-embedded gene regulation","display_name":"Brassinosteroid receptors reveal conserved features of exon-embedded gene regulation","publication_year":2026,"publication_date":"2026-09-23","ids":{"openalex":"https://openalex.org/W7214123724","doi":"https://doi.org/10.64898/2026.09.21.753065"},"language":"en","primary_location":{"id":"doi:10.64898/2026.09.21.753065","doi":"https://doi.org/10.64898/2026.09.21.753065","is_oa":true,"landing_page_url":"https://doi.org/10.64898/2026.09.21.753065","pdf_url":null,"source":{"id":"https://openalex.org/S4306402567","display_name":"bioRxiv (Cold Spring Harbor Laboratory)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I2750212522","host_organization_name":"Cold Spring Harbor Laboratory","host_organization_lineage":["https://openalex.org/I2750212522"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by-nc-nd","license_id":"https://openalex.org/licenses/cc-by-nc-nd","version":"acceptedVersion","is_accepted":true,"is_published":false,"raw_source_name":"bioRxiv","raw_type":"posted-content"},"volume":null,"type":"preprint","is_traditional":true,"indexed_in":["crossref"],"open_access":{"is_oa":true,"oa_status":"green","oa_url":"https://doi.org/10.64898/2026.09.21.753065","any_repository_has_fulltext":true},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5152557192","display_name":"Raul Denia-Mondejar","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I134820265","display_name":"Consejo Superior de Investigaciones Cient\u00edficas","ror":"https://ror.org/02gfc7t72","country_code":"ES","type":"government","lineage":["https://openalex.org/I134820265"]},{"id":"https://openalex.org/I4210164699","display_name":"Instituto de Biolog\u00eda Molecular y Celular de Plantas","ror":"https://ror.org/04zdays56","country_code":"ES","type":"facility","lineage":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]},{"id":"https://openalex.org/I60053951","display_name":"Universitat Polit\u00e8cnica de Val\u00e8ncia","ror":"https://ror.org/01460j859","country_code":"ES","type":"education","lineage":["https://openalex.org/I60053951"]}],"countries":["ES"],"is_corresponding":false,"raw_author_name":"Ra\u00fal Denia-Mond\u00e9jar","raw_affiliation_strings":["Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain","institution_ids":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5107106180","display_name":"Paula Brunot-Garau","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I134820265","display_name":"Consejo Superior de Investigaciones Cient\u00edficas","ror":"https://ror.org/02gfc7t72","country_code":"ES","type":"government","lineage":["https://openalex.org/I134820265"]},{"id":"https://openalex.org/I4210164699","display_name":"Instituto de Biolog\u00eda Molecular y Celular de Plantas","ror":"https://ror.org/04zdays56","country_code":"ES","type":"facility","lineage":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]},{"id":"https://openalex.org/I60053951","display_name":"Universitat Polit\u00e8cnica de Val\u00e8ncia","ror":"https://ror.org/01460j859","country_code":"ES","type":"education","lineage":["https://openalex.org/I60053951"]}],"countries":["ES"],"is_corresponding":false,"raw_author_name":"Paula Brunot-Garau","raw_affiliation_strings":["Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain","institution_ids":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5101899138","display_name":"Xing Ma","orcid":"https://orcid.org/0000-0001-6914-0173","observed_orcids":["https://orcid.org/0000-0001-6914-0173","https://orcid.org/0009-0004-4062-0797"]},"institutions":[{"id":"https://openalex.org/I1294671590","display_name":"Centre National de la Recherche Scientifique","ror":"https://ror.org/02feahw73","country_code":"FR","type":"government","lineage":["https://openalex.org/I1294671590"]},{"id":"https://openalex.org/I277688954","display_name":"Universit\u00e9 Paris-Saclay","ror":"https://ror.org/03xjwb503","country_code":"FR","type":"education","lineage":["https://openalex.org/I277688954"]},{"id":"https://openalex.org/I4210088668","display_name":"Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement","ror":"https://ror.org/003vg9w96","country_code":"FR","type":"government","lineage":["https://openalex.org/I4210088668"]},{"id":"https://openalex.org/I4210090571","display_name":"Institut des Sciences des Plantes de Paris Saclay","ror":"https://ror.org/00ajjta07","country_code":"FR","type":"facility","lineage":["https://openalex.org/I1294671590","https://openalex.org/I204730241","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571","https://openalex.org/I88467170"]}],"countries":["FR"],"is_corresponding":false,"raw_author_name":"Xing Ma","raw_affiliation_strings":["Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France","institution_ids":["https://openalex.org/I1294671590","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5080089793","display_name":"Antonio Serrano-Mislata","orcid":"https://orcid.org/0000-0002-8828-1809","observed_orcids":["https://orcid.org/0000-0002-8828-1809"]},"institutions":[{"id":"https://openalex.org/I134820265","display_name":"Consejo Superior de Investigaciones Cient\u00edficas","ror":"https://ror.org/02gfc7t72","country_code":"ES","type":"government","lineage":["https://openalex.org/I134820265"]},{"id":"https://openalex.org/I4210164699","display_name":"Instituto de Biolog\u00eda Molecular y Celular de Plantas","ror":"https://ror.org/04zdays56","country_code":"ES","type":"facility","lineage":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]},{"id":"https://openalex.org/I60053951","display_name":"Universitat Polit\u00e8cnica de Val\u00e8ncia","ror":"https://ror.org/01460j859","country_code":"ES","type":"education","lineage":["https://openalex.org/I60053951"]}],"countries":["ES"],"is_corresponding":false,"raw_author_name":"Antonio Serrano-Mislata","raw_affiliation_strings":["Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain"],"raw_orcid":"https://orcid.org/0000-0002-8828-1809","affiliations":[{"raw_affiliation_string":"Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain","institution_ids":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5080264896","display_name":"Mar\u00eda Dolores G\u00f3mez","orcid":"https://orcid.org/0000-0001-9445-2056","observed_orcids":["https://orcid.org/0000-0001-9445-2056"]},"institutions":[{"id":"https://openalex.org/I134820265","display_name":"Consejo Superior de Investigaciones Cient\u00edficas","ror":"https://ror.org/02gfc7t72","country_code":"ES","type":"government","lineage":["https://openalex.org/I134820265"]},{"id":"https://openalex.org/I4210164699","display_name":"Instituto de Biolog\u00eda Molecular y Celular de Plantas","ror":"https://ror.org/04zdays56","country_code":"ES","type":"facility","lineage":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]},{"id":"https://openalex.org/I60053951","display_name":"Universitat Polit\u00e8cnica de Val\u00e8ncia","ror":"https://ror.org/01460j859","country_code":"ES","type":"education","lineage":["https://openalex.org/I60053951"]}],"countries":["ES"],"is_corresponding":false,"raw_author_name":"Mar\u00eda Dolores G\u00f3mez","raw_affiliation_strings":["Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain","institution_ids":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5039000924","display_name":"David Latrasse","orcid":"https://orcid.org/0000-0003-2149-0128","observed_orcids":["https://orcid.org/0000-0003-2149-0128"]},"institutions":[{"id":"https://openalex.org/I1294671590","display_name":"Centre National de la Recherche Scientifique","ror":"https://ror.org/02feahw73","country_code":"FR","type":"government","lineage":["https://openalex.org/I1294671590"]},{"id":"https://openalex.org/I277688954","display_name":"Universit\u00e9 Paris-Saclay","ror":"https://ror.org/03xjwb503","country_code":"FR","type":"education","lineage":["https://openalex.org/I277688954"]},{"id":"https://openalex.org/I4210088668","display_name":"Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement","ror":"https://ror.org/003vg9w96","country_code":"FR","type":"government","lineage":["https://openalex.org/I4210088668"]},{"id":"https://openalex.org/I4210090571","display_name":"Institut des Sciences des Plantes de Paris Saclay","ror":"https://ror.org/00ajjta07","country_code":"FR","type":"facility","lineage":["https://openalex.org/I1294671590","https://openalex.org/I204730241","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571","https://openalex.org/I88467170"]}],"countries":["FR"],"is_corresponding":false,"raw_author_name":"David Latrasse","raw_affiliation_strings":["Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France","institution_ids":["https://openalex.org/I1294671590","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5070486620","display_name":"Moussa Benhamed","orcid":"https://orcid.org/0000-0002-2716-748X","observed_orcids":["https://orcid.org/0000-0002-2716-748X","https://orcid.org/0000-0002-4181-1702"]},"institutions":[{"id":"https://openalex.org/I1294671590","display_name":"Centre National de la Recherche Scientifique","ror":"https://ror.org/02feahw73","country_code":"FR","type":"government","lineage":["https://openalex.org/I1294671590"]},{"id":"https://openalex.org/I277688954","display_name":"Universit\u00e9 Paris-Saclay","ror":"https://ror.org/03xjwb503","country_code":"FR","type":"education","lineage":["https://openalex.org/I277688954"]},{"id":"https://openalex.org/I4210088668","display_name":"Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement","ror":"https://ror.org/003vg9w96","country_code":"FR","type":"government","lineage":["https://openalex.org/I4210088668"]},{"id":"https://openalex.org/I4210090571","display_name":"Institut des Sciences des Plantes de Paris Saclay","ror":"https://ror.org/00ajjta07","country_code":"FR","type":"facility","lineage":["https://openalex.org/I1294671590","https://openalex.org/I204730241","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571","https://openalex.org/I88467170"]}],"countries":["FR"],"is_corresponding":false,"raw_author_name":"Moussa Benhamed","raw_affiliation_strings":["Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Universit\u00e9 Paris-Saclay, CNRS, INRAE, Univ Evry, Institute of Plant Sciences Paris-Saclay (IPS2), Orsay 91405, France","institution_ids":["https://openalex.org/I1294671590","https://openalex.org/I277688954","https://openalex.org/I4210088668","https://openalex.org/I4210090571"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5053746432","display_name":"Christian S. Hardtke","orcid":"https://orcid.org/0000-0003-3203-1058","observed_orcids":["https://orcid.org/0000-0003-3203-1058"]},"institutions":[{"id":"https://openalex.org/I97565354","display_name":"University of Lausanne","ror":"https://ror.org/019whta54","country_code":"CH","type":"education","lineage":["https://openalex.org/I97565354"]}],"countries":["CH"],"is_corresponding":true,"raw_author_name":"Christian S. Hardtke","raw_affiliation_strings":["Department of Plant Molecular Biology, Biophore, UNIL-Sorge, University of Lausanne, CH-1015 Lausanne, Switzerland"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Department of Plant Molecular Biology, Biophore, UNIL-Sorge, University of Lausanne, CH-1015 Lausanne, Switzerland","institution_ids":["https://openalex.org/I97565354"]}]},{"author_position":"last","author":{"id":"https://openalex.org/A5070093006","display_name":"Noel Blanco\u2010Touri\u00f1\u00e1n","orcid":"https://orcid.org/0000-0003-4610-6110","observed_orcids":["https://orcid.org/0000-0003-4610-6110"]},"institutions":[{"id":"https://openalex.org/I134820265","display_name":"Consejo Superior de Investigaciones Cient\u00edficas","ror":"https://ror.org/02gfc7t72","country_code":"ES","type":"government","lineage":["https://openalex.org/I134820265"]},{"id":"https://openalex.org/I4210164699","display_name":"Instituto de Biolog\u00eda Molecular y Celular de Plantas","ror":"https://ror.org/04zdays56","country_code":"ES","type":"facility","lineage":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]},{"id":"https://openalex.org/I60053951","display_name":"Universitat Polit\u00e8cnica de Val\u00e8ncia","ror":"https://ror.org/01460j859","country_code":"ES","type":"education","lineage":["https://openalex.org/I60053951"]},{"id":"https://openalex.org/I97565354","display_name":"University of Lausanne","ror":"https://ror.org/019whta54","country_code":"CH","type":"education","lineage":["https://openalex.org/I97565354"]}],"countries":["CH","ES"],"is_corresponding":false,"raw_author_name":"Noel Blanco-Touri\u00f1\u00e1n","raw_affiliation_strings":["Department of Plant Molecular Biology, Biophore, UNIL-Sorge, University of Lausanne, CH-1015 Lausanne, Switzerland","Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Department of Plant Molecular Biology, Biophore, UNIL-Sorge, University of Lausanne, CH-1015 Lausanne, Switzerland","institution_ids":["https://openalex.org/I97565354"]},{"raw_affiliation_string":"Instituto de Biolog\u00eda Molecular y Celular de Plantas, Consejo Superior de Investigaciones Cient\u00edficas - Universitat Polit\u00e8cnica de Val\u00e8ncia, 46022, Valencia, Spain","institution_ids":["https://openalex.org/I134820265","https://openalex.org/I4210164699","https://openalex.org/I60053951"]}]}],"institutions":[],"countries_distinct_count":3,"institutions_distinct_count":8,"corresponding_author_ids":["https://openalex.org/A5053746432"],"corresponding_institution_ids":["https://openalex.org/I97565354"],"apc_list":null,"apc_paid":null,"fwci":null,"has_fulltext":false,"cited_by_count":0,"citation_normalized_percentile":null,"cited_by_percentile_year":null,"biblio":{"volume":null,"issue":null,"first_page":null,"last_page":null},"is_retracted":false,"is_paratext":false,"is_xpac":false,"primary_topic":{"id":"https://openalex.org/T10184","display_name":"Plant Molecular Biology Research","score":0.961899995803833,"subfield":{"id":"https://openalex.org/subfields/1110","display_name":"Plant Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},"topics":[{"id":"https://openalex.org/T10184","display_name":"Plant Molecular Biology Research","score":0.961899995803833,"subfield":{"id":"https://openalex.org/subfields/1110","display_name":"Plant Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T10222","display_name":"Genomics and Chromatin Dynamics","score":0.02019999921321869,"subfield":{"id":"https://openalex.org/subfields/1312","display_name":"Molecular Biology"},"field":{"id":"https://openalex.org/fields/13","display_name":"Biochemistry, Genetics and Molecular Biology"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T10434","display_name":"Chromosomal and Genetic Variations","score":0.010200000368058681,"subfield":{"id":"https://openalex.org/subfields/1311","display_name":"Genetics"},"field":{"id":"https://openalex.org/fields/13","display_name":"Biochemistry, Genetics and Molecular Biology"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[{"id":"https://openalex.org/keywords/bri1","display_name":"BRI1","score":0.8169999718666077},{"id":"https://openalex.org/keywords/brassinosteroid-receptor","display_name":"brassinosteroid receptor","score":0.9340000152587891},{"id":"https://openalex.org/keywords/receptor-kinases","display_name":"receptor kinases","score":0.7400000095367432},{"id":"https://openalex.org/keywords/spatial-gene-expression","display_name":"spatial gene expression","score":0.652999997138977},{"id":"https://openalex.org/keywords/root-growth","display_name":"root growth","score":0.8740000128746033},{"id":"https://openalex.org/keywords/cis-regulatory-elements","display_name":"cis-regulatory elements","score":0.8420000076293945},{"id":"https://openalex.org/keywords/chromatin-interaction-networks","display_name":"chromatin interaction networks","score":0.8539999723434448},{"id":"https://openalex.org/keywords/plant-gene-regulation","display_name":"plant gene regulation","score":0.7160000205039978},{"id":"https://openalex.org/keywords/evolution-of-gene-regulation","display_name":"gene regulatory evolution","score":0.5600000023841858}],"concepts":[{"id":"https://openalex.org/C2778778573","wikidata":"https://www.wikidata.org/wiki/Q421976","display_name":"Brassinosteroid","level":5,"score":0.8528000116348267},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.7145000100135803},{"id":"https://openalex.org/C104317684","wikidata":"https://www.wikidata.org/wiki/Q7187","display_name":"Gene","level":2,"score":0.6129000186920166},{"id":"https://openalex.org/C165864922","wikidata":"https://www.wikidata.org/wiki/Q411391","display_name":"Regulation of gene expression","level":3,"score":0.6111000180244446},{"id":"https://openalex.org/C111936080","wikidata":"https://www.wikidata.org/wiki/Q913367","display_name":"Enhancer","level":4,"score":0.5974000096321106},{"id":"https://openalex.org/C150194340","wikidata":"https://www.wikidata.org/wiki/Q26972","display_name":"Gene expression","level":3,"score":0.5859000086784363},{"id":"https://openalex.org/C54355233","wikidata":"https://www.wikidata.org/wiki/Q7162","display_name":"Genetics","level":1,"score":0.5396999716758728},{"id":"https://openalex.org/C83640560","wikidata":"https://www.wikidata.org/wiki/Q180951","display_name":"Chromatin","level":3,"score":0.5343999862670898},{"id":"https://openalex.org/C21592294","wikidata":"https://www.wikidata.org/wiki/Q3238407","display_name":"Regulatory sequence","level":4,"score":0.4715999960899353},{"id":"https://openalex.org/C33498276","wikidata":"https://www.wikidata.org/wiki/Q2138797","display_name":"Regulator gene","level":4,"score":0.46639999747276306},{"id":"https://openalex.org/C70721500","wikidata":"https://www.wikidata.org/wiki/Q177005","display_name":"Computational biology","level":1,"score":0.4530999958515167},{"id":"https://openalex.org/C84597430","wikidata":"https://www.wikidata.org/wiki/Q106227","display_name":"Locus (genetics)","level":3,"score":0.4196999967098236},{"id":"https://openalex.org/C67339327","wikidata":"https://www.wikidata.org/wiki/Q1502576","display_name":"Gene regulatory network","level":4,"score":0.412200003862381},{"id":"https://openalex.org/C27153228","wikidata":"https://www.wikidata.org/wiki/Q3787274","display_name":"Transcriptional regulation","level":4,"score":0.3799999952316284},{"id":"https://openalex.org/C190796033","wikidata":"https://www.wikidata.org/wiki/Q1093165","display_name":"Cis-regulatory module","level":5,"score":0.35850000381469727},{"id":"https://openalex.org/C101762097","wikidata":"https://www.wikidata.org/wiki/Q224093","display_name":"Promoter","level":4,"score":0.35510000586509705},{"id":"https://openalex.org/C2779491563","wikidata":"https://www.wikidata.org/wiki/Q157892","display_name":"Arabidopsis","level":4,"score":0.3379000127315521},{"id":"https://openalex.org/C62361671","wikidata":"https://www.wikidata.org/wiki/Q1754612","display_name":"Regulon","level":4,"score":0.335099995136261},{"id":"https://openalex.org/C95444343","wikidata":"https://www.wikidata.org/wiki/Q7141","display_name":"Cell biology","level":1,"score":0.33079999685287476},{"id":"https://openalex.org/C86339819","wikidata":"https://www.wikidata.org/wiki/Q407384","display_name":"Transcription factor","level":3,"score":0.3077000081539154},{"id":"https://openalex.org/C6929976","wikidata":"https://www.wikidata.org/wiki/Q3771881","display_name":"Regulator","level":3,"score":0.2985999882221222},{"id":"https://openalex.org/C62478195","wikidata":"https://www.wikidata.org/wiki/Q828130","display_name":"Signal transduction","level":2,"score":0.2687999904155731},{"id":"https://openalex.org/C18431079","wikidata":"https://www.wikidata.org/wiki/Q1502169","display_name":"Gene expression profiling","level":4,"score":0.2623000144958496},{"id":"https://openalex.org/C179926584","wikidata":"https://www.wikidata.org/wiki/Q207714","display_name":"Transcription (linguistics)","level":2,"score":0.2558000087738037}],"mesh":[],"locations_count":1,"locations":[{"id":"doi:10.64898/2026.09.21.753065","doi":"https://doi.org/10.64898/2026.09.21.753065","is_oa":true,"landing_page_url":"https://doi.org/10.64898/2026.09.21.753065","pdf_url":null,"source":{"id":"https://openalex.org/S4306402567","display_name":"bioRxiv (Cold Spring Harbor Laboratory)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I2750212522","host_organization_name":"Cold Spring Harbor Laboratory","host_organization_lineage":["https://openalex.org/I2750212522"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by-nc-nd","license_id":"https://openalex.org/licenses/cc-by-nc-nd","version":"acceptedVersion","is_accepted":true,"is_published":false,"raw_source_name":"bioRxiv","raw_type":"posted-content"}],"best_oa_location":{"id":"doi:10.64898/2026.09.21.753065","doi":"https://doi.org/10.64898/2026.09.21.753065","is_oa":true,"landing_page_url":"https://doi.org/10.64898/2026.09.21.753065","pdf_url":null,"source":{"id":"https://openalex.org/S4306402567","display_name":"bioRxiv (Cold Spring Harbor Laboratory)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I2750212522","host_organization_name":"Cold Spring Harbor Laboratory","host_organization_lineage":["https://openalex.org/I2750212522"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by-nc-nd","license_id":"https://openalex.org/licenses/cc-by-nc-nd","version":"acceptedVersion","is_accepted":true,"is_published":false,"raw_source_name":"bioRxiv","raw_type":"posted-content"},"sustainable_development_goals":[],"study_designs":[],"awards":[],"funders":[],"has_content":{"pdf":false,"grobid_xml":false},"content_urls":null,"referenced_works_count":68,"referenced_works":["https://openalex.org/W1905057662","https://openalex.org/W1969353942","https://openalex.org/W1977214336","https://openalex.org/W1982196595","https://openalex.org/W1996497687","https://openalex.org/W1999143685","https://openalex.org/W2002115219","https://openalex.org/W2014677321","https://openalex.org/W2021341670","https://openalex.org/W2023849539","https://openalex.org/W2031310524","https://openalex.org/W2059979506","https://openalex.org/W2075272534","https://openalex.org/W2096315332","https://openalex.org/W2106882534","https://openalex.org/W2107337799","https://openalex.org/W2114280998","https://openalex.org/W2121150295","https://openalex.org/W2136430672","https://openalex.org/W2140952049","https://openalex.org/W2141292707","https://openalex.org/W2142320568","https://openalex.org/W2144775551","https://openalex.org/W2151763226","https://openalex.org/W2157009395","https://openalex.org/W2157294409","https://openalex.org/W2167279371","https://openalex.org/W2170551349","https://openalex.org/W2201264108","https://openalex.org/W2264491011","https://openalex.org/W2462790555","https://openalex.org/W2533355719","https://openalex.org/W2568479992","https://openalex.org/W2575118365","https://openalex.org/W2793057599","https://openalex.org/W2902291530","https://openalex.org/W2963423318","https://openalex.org/W2983263940","https://openalex.org/W2988396470","https://openalex.org/W2995638580","https://openalex.org/W3013689061","https://openalex.org/W3037009539","https://openalex.org/W3049718544","https://openalex.org/W3131055609","https://openalex.org/W3134968252","https://openalex.org/W3135435890","https://openalex.org/W3189548749","https://openalex.org/W4206011972","https://openalex.org/W4284990903","https://openalex.org/W4289835134","https://openalex.org/W4313611824","https://openalex.org/W4318340527","https://openalex.org/W4382631713","https://openalex.org/W4392157945","https://openalex.org/W4394785938","https://openalex.org/W4400352120","https://openalex.org/W4400780205","https://openalex.org/W4401641305","https://openalex.org/W4402830861","https://openalex.org/W4404511000","https://openalex.org/W4405670661","https://openalex.org/W4409538612","https://openalex.org/W4415970773","https://openalex.org/W4416263400","https://openalex.org/W4416863496","https://openalex.org/W4416925580","https://openalex.org/W7147689203","https://openalex.org/W7161757636"],"related_works":["https://openalex.org/W2097348290","https://openalex.org/W2039084633","https://openalex.org/W7147689203","https://openalex.org/W2062612066","https://openalex.org/W2951735260","https://openalex.org/W7098973182","https://openalex.org/W2333457370","https://openalex.org/W3048321509","https://openalex.org/W3148293239","https://openalex.org/W1558369695"],"abstract_inverted_index":{"Abstract":[0],"Precise":[1],"quantitative":[2],"and":[3,18,33,48,72,81,92,119],"spatial":[4,63],"control":[5],"of":[6,130,147],"plant":[7,30,137,148],"gene":[8,31,149],"expression":[9,32,65,71],"is":[10,58,143],"typically":[11],"attributed":[12],"to":[13,24,60],"cis-regulatory":[14],"elements":[15],"in":[16,87],"promoters":[17],"other":[19],"non-coding":[20],"sequences.":[21,104],"The":[22,83],"extent":[23],"which":[25,57],"exon-encoded":[26,78,131,140],"regulatory":[27,79,99,132,141],"information":[28,80,100,133,142],"shapes":[29],"how":[34],"remains":[35],"unclear.":[36],"Here":[37],"we":[38],"used":[39],"BRASSINOSTEROID":[40],"INSENSITIVE":[41],"1":[42],"(BRI1)":[43],"as":[44],"a":[45,144],"model":[46],"locus":[47],"found":[49],"that":[50,108,127],"it":[51],"contains":[52],"an":[53],"exonic":[54],"enhancer":[55],"(EE),":[56],"essential":[59],"establish":[61],"the":[62],"BRI1":[64,70,84],"pattern.":[66],"Its":[67],"disruption":[68],"restricts":[69],"impairs":[73],"BRI1-dependent":[74],"root":[75],"growth,":[76],"linking":[77],"development.":[82],"EE":[85],"acts":[86],"concert":[88],"with":[89,113],"its":[90],"promoter,":[91],"related":[93],"receptor":[94],"kinase":[95],"genes":[96],"likewise":[97],"harbor":[98],"within":[101],"their":[102],"coding":[103],"Genome-wide":[105],"surveys":[106],"show":[107],"candidate":[109],"EEs":[110],"are":[111],"associated":[112],"highly":[114],"connected":[115],"chromatin":[116],"interaction":[117],"networks":[118],"transcriptionally":[120],"active":[121],"loci.":[122],"Moreover,":[123],"species":[124],"comparisons":[125],"suggest":[126],"common":[128],"features":[129],"extend":[134],"across":[135],"diverse":[136],"lineages.":[138],"Thus,":[139],"vital":[145],"component":[146],"regulation.":[150]},"counts_by_year":[],"updated_date":"2026-11-08T12:12:28.789561","created_date":"2026-09-24T00:00:00"}