{"id":"https://openalex.org/W7212157366","doi":"https://doi.org/10.15468/tn5xar","title":"A systematic review of the Limnonectes hascheanus-limborgi complex (Anura: Dicroglossidae), with the description of a new species from Peninsular Malaysia","display_name":"A systematic review of the Limnonectes hascheanus-limborgi complex (Anura: Dicroglossidae), with the description of a new species from Peninsular Malaysia","publication_year":2026,"publication_date":"2026-09-10","ids":{"openalex":"https://openalex.org/W7212157366","doi":"https://doi.org/10.15468/tn5xar"},"language":"en","primary_location":{"id":"doi:10.15468/tn5xar","doi":"https://doi.org/10.15468/tn5xar","is_oa":true,"landing_page_url":"https://doi.org/10.15468/tn5xar","pdf_url":null,"source":{"id":"https://openalex.org/S4398183272","display_name":"Global Biodiversity Information Facility","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I111796040","host_organization_name":"Global Biodiversity Information Facility","host_organization_lineage":["https://openalex.org/I111796040"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":null,"is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Dataset"},"volume":null,"type":"dataset","is_traditional":false,"indexed_in":["datacite"],"open_access":{"is_oa":true,"oa_status":"green","oa_url":"https://doi.org/10.15468/tn5xar","any_repository_has_fulltext":true},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5024445084","display_name":"Bei Zou","orcid":"https://orcid.org/0000-0001-8480-1419","observed_orcids":["https://orcid.org/0000-0001-8480-1419"]},"institutions":[{"id":"https://openalex.org/I139322472","display_name":"Universiti Sains Malaysia","ror":"https://ror.org/02rgb2k63","country_code":"MY","type":"education","lineage":["https://openalex.org/I139322472"]}],"countries":["MY"],"is_corresponding":false,"raw_author_name":"Bei Zou","raw_affiliation_strings":["School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia","institution_ids":["https://openalex.org/I139322472"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5123228859","display_name":"M. S. Shahrul Anuar","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I139322472","display_name":"Universiti Sains Malaysia","ror":"https://ror.org/02rgb2k63","country_code":"MY","type":"education","lineage":["https://openalex.org/I139322472"]}],"countries":["MY"],"is_corresponding":false,"raw_author_name":"M. S. Shahrul Anuar","raw_affiliation_strings":["School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia. & Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia. & Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia","institution_ids":["https://openalex.org/I139322472"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5095850987","display_name":"TEK JOO LOW","orcid":"https://orcid.org/0009-0005-9883-9027","observed_orcids":["https://orcid.org/0009-0005-9883-9027"]},"institutions":[{"id":"https://openalex.org/I139322472","display_name":"Universiti Sains Malaysia","ror":"https://ror.org/02rgb2k63","country_code":"MY","type":"education","lineage":["https://openalex.org/I139322472"]}],"countries":["MY"],"is_corresponding":false,"raw_author_name":"Tek Joo Low","raw_affiliation_strings":["School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia","institution_ids":["https://openalex.org/I139322472"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5044076954","display_name":"ZIJIA HONG","orcid":"https://orcid.org/0009-0005-4554-1315","observed_orcids":["https://orcid.org/0009-0005-4554-1315"]},"institutions":[{"id":"https://openalex.org/I139322472","display_name":"Universiti Sains Malaysia","ror":"https://ror.org/02rgb2k63","country_code":"MY","type":"education","lineage":["https://openalex.org/I139322472"]}],"countries":["MY"],"is_corresponding":false,"raw_author_name":"Zijia Hong","raw_affiliation_strings":["School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Penang, Malaysia","institution_ids":["https://openalex.org/I139322472"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5139899189","display_name":"Elizabeth Chau","orcid":"https://orcid.org/0009-0006-9417-6200","observed_orcids":["https://orcid.org/0009-0006-9417-6200"]},"institutions":[],"countries":[],"is_corresponding":false,"raw_author_name":"Elizabeth Chau","raw_affiliation_strings":[],"raw_orcid":"https://orcid.org/0000-0001-8422-3698","affiliations":[]},{"author_position":"middle","author":{"id":"https://openalex.org/A5121095595","display_name":"L Lee Grismer","orcid":"https://orcid.org/0000-0001-8422-3698","observed_orcids":["https://orcid.org/0000-0001-8422-3698"]},"institutions":[{"id":"https://openalex.org/I2801277401","display_name":"San Diego Natural History Museum","ror":"https://ror.org/00kmpab62","country_code":"US","type":"archive","lineage":["https://openalex.org/I2801277401"]}],"countries":["US"],"is_corresponding":false,"raw_author_name":"L. Lee Grismer","raw_affiliation_strings":["Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA. & El Serpentario y C. E. M. A de Baja California Sur, Brercha California e / Nuevo Reforma y Guaycura. Fracc: Benito Ju\u00e1rez, CP 23090, La Paz, Baja California Sur, M\u00e9xico"],"raw_orcid":"https://orcid.org/0000-0001-8422-3698","affiliations":[{"raw_affiliation_string":"Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA. & El Serpentario y C. E. M. A de Baja California Sur, Brercha California e / Nuevo Reforma y Guaycura. Fracc: Benito Ju\u00e1rez, CP 23090, La Paz, Baja California Sur, M\u00e9xico","institution_ids":["https://openalex.org/I2801277401"]}]},{"author_position":"last","author":{"id":"https://openalex.org/A5067421892","display_name":"Evan S. H. Quah","orcid":"https://orcid.org/0000-0002-5357-1953","observed_orcids":["https://orcid.org/0000-0002-5357-1953"]},"institutions":[{"id":"https://openalex.org/I165932596","display_name":"National University of Singapore","ror":"https://ror.org/01tgyzw49","country_code":"SG","type":"education","lineage":["https://openalex.org/I165932596"]}],"countries":["SG"],"is_corresponding":false,"raw_author_name":"Evan S. H. Quah","raw_affiliation_strings":["Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377"],"raw_orcid":"https://orcid.org/0000-0002-5357-1953","affiliations":[{"raw_affiliation_string":"Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377","institution_ids":["https://openalex.org/I165932596"]}]}],"institutions":[],"countries_distinct_count":3,"institutions_distinct_count":3,"corresponding_author_ids":[],"corresponding_institution_ids":[],"apc_list":null,"apc_paid":null,"fwci":null,"has_fulltext":false,"cited_by_count":0,"citation_normalized_percentile":null,"cited_by_percentile_year":null,"biblio":{"volume":null,"issue":null,"first_page":null,"last_page":null},"is_retracted":false,"is_paratext":false,"is_xpac":false,"primary_topic":{"id":"https://openalex.org/T10332","display_name":"Amphibian and Reptile Biology","score":0.9951000213623047,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},"topics":[{"id":"https://openalex.org/T10332","display_name":"Amphibian and Reptile Biology","score":0.9951000213623047,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T12319","display_name":"Fish Biology and Ecology Studies","score":0.0012000000569969416,"subfield":{"id":"https://openalex.org/subfields/1104","display_name":"Aquatic Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T13269","display_name":"Orthoptera Research and Taxonomy","score":0.000699999975040555,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[],"concepts":[{"id":"https://openalex.org/C203358219","wikidata":"https://www.wikidata.org/wiki/Q223209","display_name":"Tympanum (architecture)","level":5,"score":0.8483999967575073},{"id":"https://openalex.org/C2781140086","wikidata":"https://www.wikidata.org/wiki/Q557945","display_name":"Confusion","level":2,"score":0.5602999925613403},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.4462999999523163},{"id":"https://openalex.org/C125390029","wikidata":"https://www.wikidata.org/wiki/Q1297859","display_name":"Species complex","level":4,"score":0.4239000082015991},{"id":"https://openalex.org/C3017739461","wikidata":"https://www.wikidata.org/wiki/Q40285","display_name":"New guinea","level":2,"score":0.40450000762939453},{"id":"https://openalex.org/C58642233","wikidata":"https://www.wikidata.org/wiki/Q8269924","display_name":"Taxonomy (biology)","level":2,"score":0.3711000084877014},{"id":"https://openalex.org/C78458016","wikidata":"https://www.wikidata.org/wiki/Q840400","display_name":"Evolutionary biology","level":1,"score":0.35530000925064087},{"id":"https://openalex.org/C90856448","wikidata":"https://www.wikidata.org/wiki/Q431","display_name":"Zoology","level":1,"score":0.3280999958515167},{"id":"https://openalex.org/C105702510","wikidata":"https://www.wikidata.org/wiki/Q514","display_name":"Anatomy","level":1,"score":0.3084000051021576},{"id":"https://openalex.org/C205649164","wikidata":"https://www.wikidata.org/wiki/Q1071","display_name":"Geography","level":0,"score":0.27250000834465027},{"id":"https://openalex.org/C164866538","wikidata":"https://www.wikidata.org/wiki/Q367351","display_name":"Cluster (spacecraft)","level":2,"score":0.2687000036239624}],"mesh":[],"locations_count":1,"locations":[{"id":"doi:10.15468/tn5xar","doi":"https://doi.org/10.15468/tn5xar","is_oa":true,"landing_page_url":"https://doi.org/10.15468/tn5xar","pdf_url":null,"source":{"id":"https://openalex.org/S4398183272","display_name":"Global Biodiversity Information Facility","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I111796040","host_organization_name":"Global Biodiversity Information Facility","host_organization_lineage":["https://openalex.org/I111796040"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":null,"is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Dataset"}],"best_oa_location":{"id":"doi:10.15468/tn5xar","doi":"https://doi.org/10.15468/tn5xar","is_oa":true,"landing_page_url":"https://doi.org/10.15468/tn5xar","pdf_url":null,"source":{"id":"https://openalex.org/S4398183272","display_name":"Global Biodiversity Information Facility","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I111796040","host_organization_name":"Global Biodiversity Information Facility","host_organization_lineage":["https://openalex.org/I111796040"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":null,"is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Dataset"},"sustainable_development_goals":[],"study_designs":[],"awards":[],"funders":[],"has_content":{"pdf":false,"grobid_xml":false},"content_urls":null,"referenced_works_count":0,"referenced_works":[],"related_works":["https://openalex.org/W4390690707","https://openalex.org/W7166418484","https://openalex.org/W6929974592","https://openalex.org/W7203654657","https://openalex.org/W6943376316","https://openalex.org/W6906398217","https://openalex.org/W6943593347","https://openalex.org/W2155453391","https://openalex.org/W6893600668","https://openalex.org/W6893559726"],"abstract_inverted_index":{"This":[0],"dataset":[1],"contains":[2],"the":[3,10,39,46,68,83,91,107,127,164,169,210,218,261,264,271,286,357,370,373,378,385,396],"digitized":[4],"treatments":[5],"in":[6,72,90,97,115,149,181,186,335,351,382,384,395],"Plazi":[7],"based":[8,93,162],"on":[9,94,163,233],"original":[11],"journal":[12],"article":[13],"Zou,":[14],"Bei,":[15],"Anuar,":[16],"M.":[17],"S.":[18,32],"Shahrul,":[19],"Low,":[20],"Tek":[21],"Joo,":[22],"Hong,":[23],"Zijia,":[24],"Chau,":[25],"Elizabeth,":[26],"Grismer,":[27],"L.":[28,86,101,111,147,349,360],"Lee,":[29],"Quah,":[30],"Evan":[31],"H.":[33],"(2026):":[34],"A":[35],"systematic":[36],"review":[37],"of":[38,48,67,85,110,129,160,209,230,260,263,285,348,356,372],"Limnonectes":[40,69,133,140],"hascheanus-limborgi":[41,70],"complex":[42,71,113],"(Anura:":[43],"Dicroglossidae),":[44],"with":[45,146,177,240,278,289],"description":[47],"a":[49,80,130],"new":[50,131,358],"species":[51,109,359],"from":[52,137],"Peninsular":[53,73,116,138,352],"Malaysia.":[54,117,139],"Zootaxa":[55],"5785":[56],"(1):":[57],"67-102,":[58],"DOI:":[59],"10.11646/zootaxa.5785.1.3,":[60],"URL:":[61],"https://doi.org/10.11646/zootaxa.5785.1.3":[62],"Abstract":[63],"The":[64,340],"taxonomic":[65],"confusion":[66],"Malaysia":[74,353],"has":[75],"lasted":[76],"for":[77],"more":[78],"than":[79,258],"decade":[81],"since":[82],"presence":[84],"limborgi":[87,350],"was":[88,103],"reported":[89,346],"country":[92],"molecular":[95,122],"data":[96],"2014.":[98],"Before":[99],"that,":[100],"hascheanus":[102,148],"thought":[104],"to":[105,221,281,369,376],"be":[106],"only":[108],"hascheaus-limborgi":[112],"occurring":[114],"Currently,":[118],"comprehensive":[119],"approaches":[120],"using":[121],"and":[123,168,191,235,256,338,364,387,392],"morphological":[124,172],"analyses":[125],"corroborate":[126],"existence":[128],"species,":[132],"corvus":[134,141,361],"sp.":[135,142,362],"nov.":[136,143,363],"is":[144],"syntopic":[145],"some":[150],"localities":[151],"but":[152],"differs":[153],"by":[154],"having":[155],"an":[156],"average":[157],"uncorrected":[158],"p-distance":[159],"7.8%":[161],"16S":[165],"rRNA":[166],"gene":[167],"following":[170],"diagnostic":[171],"characteristics:":[173],"(1)":[174],"small":[175],"size":[176],"SVL":[178,183],"21.59\u201335.45":[179],"mm":[180,185],"males,":[182],"28.74\u201335.78":[184],"females;":[187],"(2)":[188],"short,":[189],"stocky":[190],"robust":[192],"body;":[193],"(3)":[194],"males":[195,337],"lack":[196],"external":[197],"vocal":[198],"sacs;":[199],"(4)":[200],"loreal":[201],"region":[202,237],"slightly":[203],"concave;":[204],"(5)":[205],"tympanum":[206,219],"approximately":[207],"2/3":[208],"eye":[211],"diameter;":[212],"(6)":[213],"distinct":[214],"supratympanic":[215],"fold":[216],"along":[217,270],"extending":[220,280],"supra-axillary":[222],"region;":[223],"(7)":[224],"smooth":[225],"skin":[226],"without":[227],"dense":[228],"cluster":[229],"granular":[231],"warts":[232],"super-axillary":[234],"cloacal":[236],"(8)":[238],"fingertips":[239],"terminal":[241],"bulbs;":[242],"(9)":[243],"relative":[244],"finger":[245],"lengths":[246],"III>I=II>IV":[247],"or":[248,315],"III>IV=I=II;":[249],"(10)":[250],"inner":[251],"metatarsal":[252],"tubercle":[253,284],"distinctly":[254],"raised":[255],"less":[257],"50%":[259],"length":[262],"first":[265],"toe;":[266],"(11)":[267],"dermal":[268],"ridge":[269],"outer":[272],"metatarsal;":[273],"(12)":[274],"toes":[275],"basally":[276],"webbed,":[277],"webbing":[279,290],"middle":[282],"subarticular":[283],"fourth":[287],"toe":[288],"formula":[291],"as":[292],"follow:":[293],"I":[294,316],"1":[295,306,317,321],"-":[296,318,328],"\u20132":[297,319],"II":[298,320],"0":[299],"+":[300,307,322],"\u2013":[301],"22":[302],"/":[303,312],"3":[304],"III":[305,324],"\u20133":[308],"IV":[309,326],"3\u2013":[310],"11":[311],"2":[313],"V":[314],"\u20132\u00bd":[323],"1\u00bd\u20133":[325],"3\u20132":[327],"IV;":[329],"(13)":[330],"enlarged":[331],"odontoids":[332],"are":[333,354],"present":[334],"both":[336],"females.":[339],"current":[341],"study":[342],"found":[343],"that":[344],"previously":[345],"populations":[347],"misidentifications":[355],"we":[365],"update":[366],"its":[367],"distribution":[368],"east":[371],"Bintang":[374],"range":[375],"include":[377],"Sekayu":[379],"lowland":[380],"forest":[381],"Terengganu":[383],"north":[386],"Ketumbar":[388],"Hill,":[389],"Kuala":[390],"Lumpur,":[391],"Tioman":[393],"Island":[394],"south.":[397]},"counts_by_year":[],"updated_date":"2026-10-29T09:56:32.521200","created_date":"2026-09-11T00:00:00"}