{"id":"https://openalex.org/W7166872003","doi":"https://doi.org/10.1038/s41586-026-10713-2","title":"Urokodia sheds light on the origin of chelicerae and book gills of Chelicerata","display_name":"Urokodia sheds light on the origin of chelicerae and book gills of Chelicerata","publication_year":2026,"publication_date":"2026-07-01","ids":{"openalex":"https://openalex.org/W7166872003","doi":"https://doi.org/10.1038/s41586-026-10713-2","pmid":"https://pubmed.ncbi.nlm.nih.gov/42386960"},"language":"en","primary_location":{"id":"doi:10.1038/s41586-026-10713-2","doi":"https://doi.org/10.1038/s41586-026-10713-2","is_oa":true,"landing_page_url":"https://doi.org/10.1038/s41586-026-10713-2","pdf_url":null,"source":{"id":"https://openalex.org/S137773608","display_name":"Nature","issn_l":"0028-0836","issn":["0028-0836","1476-4687"],"is_oa":false,"is_in_doaj":false,"is_core":true,"listed_in":["anvur-class-a","anvur-scientific","cwts-core","dongbi-a","doyens","erih-plus","jufo-3","ki-jl-3","medline","norway-2","poland-200","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310319908","host_organization_name":"Nature Portfolio","host_organization_lineage":["https://openalex.org/P4310319908","https://openalex.org/P4310319965"],"host_organization_lineage_names":["Nature Portfolio","Springer Nature"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Nature","raw_type":"journal-article"},"volume":null,"type":"article","indexed_in":["crossref","pubmed"],"open_access":{"is_oa":true,"oa_status":"hybrid","oa_url":"https://doi.org/10.1038/s41586-026-10713-2","any_repository_has_fulltext":true},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5047333154","display_name":"Yu Liu","orcid":"https://orcid.org/0000-0002-2346-740X","observed_orcids":["https://orcid.org/0000-0002-2346-740X"]},"institutions":[{"id":"https://openalex.org/I153648349","display_name":"University of Leicester","ror":"https://ror.org/04h699437","country_code":"GB","type":"education","lineage":["https://openalex.org/I153648349"]},{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]},{"id":"https://openalex.org/I5343935","display_name":"Guilin University of Electronic Technology","ror":"https://ror.org/05arjae42","country_code":"CN","type":"education","lineage":["https://openalex.org/I5343935"]}],"countries":["CN","GB"],"is_corresponding":true,"raw_author_name":"Yu Liu","raw_affiliation_strings":["Guilin University of Electronic Technology, Guilin, China. yu.liu@ynu.edu.cn","MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China. yu.liu@ynu.edu.cn","School of Geography, Geology and the Environment, University of Leicester, Leicester, UK. yu.liu@ynu.edu.cn","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China. yu.liu@ynu.edu.cn"],"raw_orcid":"https://orcid.org/0000-0002-2346-740X","affiliations":[{"raw_affiliation_string":"Guilin University of Electronic Technology, Guilin, China. yu.liu@ynu.edu.cn","institution_ids":["https://openalex.org/I5343935"]},{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China. yu.liu@ynu.edu.cn","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"School of Geography, Geology and the Environment, University of Leicester, Leicester, UK. yu.liu@ynu.edu.cn","institution_ids":["https://openalex.org/I153648349"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China. yu.liu@ynu.edu.cn","institution_ids":["https://openalex.org/I189210763"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5018346882","display_name":"Lorenzo Lustri","orcid":"https://orcid.org/0000-0002-8951-5160","observed_orcids":["https://orcid.org/0000-0002-8951-5160"]},"institutions":[{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Lorenzo Lustri","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5011314665","display_name":"Huijuan Mai","orcid":"https://orcid.org/0000-0002-3157-263X","observed_orcids":["https://orcid.org/0000-0002-3157-263X"]},"institutions":[{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Huijuan Mai","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5011307993","display_name":"Mark Williams","orcid":"https://orcid.org/0000-0002-7987-6069","observed_orcids":["https://orcid.org/0000-0002-7987-6069"]},"institutions":[{"id":"https://openalex.org/I153648349","display_name":"University of Leicester","ror":"https://ror.org/04h699437","country_code":"GB","type":"education","lineage":["https://openalex.org/I153648349"]},{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN","GB"],"is_corresponding":false,"raw_author_name":"Mark Williams","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","School of Geography, Geology and the Environment, University of Leicester, Leicester, UK"],"raw_orcid":"https://orcid.org/0000-0002-7987-6069","affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"School of Geography, Geology and the Environment, University of Leicester, Leicester, UK","institution_ids":["https://openalex.org/I153648349"]}]},{"author_position":"middle","author":{"id":null,"display_name":"Liming Liu","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Liming Liu","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5139785793","display_name":"Yani Tang","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN"],"is_corresponding":false,"raw_author_name":"Yani Tang","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5023764195","display_name":"Thomas H. P. Harvey","orcid":"https://orcid.org/0000-0002-2717-7004","observed_orcids":["https://orcid.org/0000-0002-2717-7004"]},"institutions":[{"id":"https://openalex.org/I153648349","display_name":"University of Leicester","ror":"https://ror.org/04h699437","country_code":"GB","type":"education","lineage":["https://openalex.org/I153648349"]},{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN","GB"],"is_corresponding":false,"raw_author_name":"Thomas H. P. Harvey","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","School of Geography, Geology and the Environment, University of Leicester, Leicester, UK"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"School of Geography, Geology and the Environment, University of Leicester, Leicester, UK","institution_ids":["https://openalex.org/I153648349"]}]},{"author_position":"last","author":{"id":"https://openalex.org/A5101984090","display_name":"Xianguang Hou","orcid":"https://orcid.org/0000-0003-0634-2100","observed_orcids":["https://orcid.org/0000-0003-0634-2100"]},"institutions":[{"id":"https://openalex.org/I189210763","display_name":"Yunnan University","ror":"https://ror.org/0040axw97","country_code":"CN","type":"education","lineage":["https://openalex.org/I189210763"]}],"countries":["CN"],"is_corresponding":true,"raw_author_name":"Xianguang Hou","raw_affiliation_strings":["MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China. xghou@ynu.edu.cn","Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China. xghou@ynu.edu.cn"],"raw_orcid":"https://orcid.org/0000-0003-0634-2100","affiliations":[{"raw_affiliation_string":"MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Institute of Palaeontology, Yunnan University, Kunming, China. xghou@ynu.edu.cn","institution_ids":["https://openalex.org/I189210763"]},{"raw_affiliation_string":"Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China. xghou@ynu.edu.cn","institution_ids":["https://openalex.org/I189210763"]}]}],"institutions":[],"countries_distinct_count":2,"institutions_distinct_count":3,"corresponding_author_ids":["https://openalex.org/A5047333154","https://openalex.org/A5101984090"],"corresponding_institution_ids":["https://openalex.org/I153648349","https://openalex.org/I189210763","https://openalex.org/I5343935"],"apc_list":{"value":12690,"currency":"USD","value_usd":12690},"apc_paid":{"value":12690,"currency":"USD","value_usd":12690},"fwci":0.0,"has_fulltext":false,"cited_by_count":0,"citation_normalized_percentile":{"value":0.4920241,"is_in_top_1_percent":false,"is_in_top_10_percent":false},"cited_by_percentile_year":null,"biblio":{"volume":"656","issue":"8128","first_page":"653","last_page":"658"},"is_retracted":false,"is_paratext":false,"is_xpac":false,"primary_topic":{"id":"https://openalex.org/T10955","display_name":"Paleontology and Evolutionary Biology","score":0.8949999809265137,"subfield":{"id":"https://openalex.org/subfields/1911","display_name":"Paleontology"},"field":{"id":"https://openalex.org/fields/19","display_name":"Earth and Planetary Sciences"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},"topics":[{"id":"https://openalex.org/T10955","display_name":"Paleontology and Evolutionary Biology","score":0.8949999809265137,"subfield":{"id":"https://openalex.org/subfields/1911","display_name":"Paleontology"},"field":{"id":"https://openalex.org/fields/19","display_name":"Earth and Planetary Sciences"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},{"id":"https://openalex.org/T10109","display_name":"Paleontology and Stratigraphy of Fossils","score":0.027499999850988388,"subfield":{"id":"https://openalex.org/subfields/1911","display_name":"Paleontology"},"field":{"id":"https://openalex.org/fields/19","display_name":"Earth and Planetary Sciences"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},{"id":"https://openalex.org/T12001","display_name":"Fossil Insects in Amber","score":0.024000000208616257,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[{"id":"https://openalex.org/keywords/chengjiang-biota","display_name":"Chengjiang biota","score":0.9470000267028809},{"id":"https://openalex.org/keywords/hypostome","display_name":"hypostome","score":0.781000018119812},{"id":"https://openalex.org/keywords/x-ray-microtomography","display_name":"X-ray microtomography","score":0.7329999804496765}],"concepts":[{"id":"https://openalex.org/C193842886","wikidata":"https://www.wikidata.org/wiki/Q498963","display_name":"Arthropod mouthparts","level":2,"score":0.7874000072479248},{"id":"https://openalex.org/C105702510","wikidata":"https://www.wikidata.org/wiki/Q514","display_name":"Anatomy","level":1,"score":0.6348000168800354},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.43070000410079956},{"id":"https://openalex.org/C71924100","wikidata":"https://www.wikidata.org/wiki/Q11190","display_name":"Medicine","level":0,"score":0.31869998574256897},{"id":"https://openalex.org/C161153951","wikidata":"https://www.wikidata.org/wiki/Q132390","display_name":"Gill","level":3,"score":0.31060001254081726},{"id":"https://openalex.org/C127313418","wikidata":"https://www.wikidata.org/wiki/Q1069","display_name":"Geology","level":0,"score":0.2069000005722046},{"id":"https://openalex.org/C30055919","wikidata":"https://www.wikidata.org/wiki/Q1566548","display_name":"Cuticle (hair)","level":2,"score":0.19699999690055847},{"id":"https://openalex.org/C2778311950","wikidata":"https://www.wikidata.org/wiki/Q7363","display_name":"Nose","level":2,"score":0.19599999487400055},{"id":"https://openalex.org/C90856448","wikidata":"https://www.wikidata.org/wiki/Q431","display_name":"Zoology","level":1,"score":0.1889999955892563},{"id":"https://openalex.org/C91762617","wikidata":"https://www.wikidata.org/wiki/Q48422","display_name":"Cadaver","level":2,"score":0.181099995970726}],"mesh":[{"descriptor_ui":"D000818","descriptor_name":"Animals","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D000818","descriptor_name":"Animals","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D000818","descriptor_name":"Animals","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":true},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":true},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":true},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000145","qualifier_name":"classification","is_major_topic":true},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000145","qualifier_name":"classification","is_major_topic":true},{"descriptor_ui":"D001181","descriptor_name":"Arthropods","qualifier_ui":"Q000145","qualifier_name":"classification","is_major_topic":true},{"descriptor_ui":"D002681","descriptor_name":"China","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D002681","descriptor_name":"China","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D002681","descriptor_name":"China","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D005075","descriptor_name":"Biological Evolution","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005075","descriptor_name":"Biological Evolution","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005075","descriptor_name":"Biological Evolution","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005121","descriptor_name":"Extremities","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D005121","descriptor_name":"Extremities","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D005121","descriptor_name":"Extremities","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D005580","descriptor_name":"Fossils","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005580","descriptor_name":"Fossils","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005580","descriptor_name":"Fossils","qualifier_ui":null,"qualifier_name":null,"is_major_topic":true},{"descriptor_ui":"D005880","descriptor_name":"Gills","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":true},{"descriptor_ui":"D005880","descriptor_name":"Gills","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D005880","descriptor_name":"Gills","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D006257","descriptor_name":"Head","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D006257","descriptor_name":"Head","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D006257","descriptor_name":"Head","qualifier_ui":"Q000033","qualifier_name":"anatomy & histology","is_major_topic":false},{"descriptor_ui":"D010802","descriptor_name":"Phylogeny","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D010802","descriptor_name":"Phylogeny","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D010802","descriptor_name":"Phylogeny","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D055114","descriptor_name":"X-Ray Microtomography","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D055114","descriptor_name":"X-Ray Microtomography","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false},{"descriptor_ui":"D055114","descriptor_name":"X-Ray Microtomography","qualifier_ui":null,"qualifier_name":null,"is_major_topic":false}],"locations_count":3,"locations":[{"id":"doi:10.1038/s41586-026-10713-2","doi":"https://doi.org/10.1038/s41586-026-10713-2","is_oa":true,"landing_page_url":"https://doi.org/10.1038/s41586-026-10713-2","pdf_url":null,"source":{"id":"https://openalex.org/S137773608","display_name":"Nature","issn_l":"0028-0836","issn":["0028-0836","1476-4687"],"is_oa":false,"is_in_doaj":false,"is_core":true,"listed_in":["anvur-class-a","anvur-scientific","cwts-core","dongbi-a","doyens","erih-plus","jufo-3","ki-jl-3","medline","norway-2","poland-200","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310319908","host_organization_name":"Nature Portfolio","host_organization_lineage":["https://openalex.org/P4310319908","https://openalex.org/P4310319965"],"host_organization_lineage_names":["Nature Portfolio","Springer Nature"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Nature","raw_type":"journal-article"},{"id":"pmid:42386960","doi":null,"is_oa":false,"landing_page_url":"https://pubmed.ncbi.nlm.nih.gov/42386960","pdf_url":null,"source":{"id":"https://openalex.org/S4306525036","display_name":"PubMed","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I1299303238","host_organization_name":"National Institutes of Health","host_organization_lineage":["https://openalex.org/I1299303238"],"host_organization_lineage_names":[],"type":"repository"},"license":null,"license_id":null,"version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Nature","raw_type":"Journal Article"},{"id":"pmh:oai:figshare.com:article/32942198","doi":null,"is_oa":true,"landing_page_url":"https://figshare.com/articles/journal_contribution/Urokodia_sheds_light_on_the_origin_of_chelicerae_and_book_gills_of_Chelicerata/32942198","pdf_url":null,"source":{"id":"https://openalex.org/S4377196282","display_name":"Figshare","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I4210132348","host_organization_name":"Figshare (United Kingdom)","host_organization_lineage":["https://openalex.org/I4210132348"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Text"}],"best_oa_location":{"id":"doi:10.1038/s41586-026-10713-2","doi":"https://doi.org/10.1038/s41586-026-10713-2","is_oa":true,"landing_page_url":"https://doi.org/10.1038/s41586-026-10713-2","pdf_url":null,"source":{"id":"https://openalex.org/S137773608","display_name":"Nature","issn_l":"0028-0836","issn":["0028-0836","1476-4687"],"is_oa":false,"is_in_doaj":false,"is_core":true,"listed_in":["anvur-class-a","anvur-scientific","cwts-core","dongbi-a","doyens","erih-plus","jufo-3","ki-jl-3","medline","norway-2","poland-200","russia-white-list-1","vabb-shw"],"host_organization":"https://openalex.org/P4310319908","host_organization_name":"Nature Portfolio","host_organization_lineage":["https://openalex.org/P4310319908","https://openalex.org/P4310319965"],"host_organization_lineage_names":["Nature Portfolio","Springer Nature"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"Nature","raw_type":"journal-article"},"sustainable_development_goals":[],"sustainable_development_goals_aurora":[],"study_designs":[],"awards":[],"funders":[],"has_content":{"grobid_xml":false,"pdf":false},"content_urls":null,"referenced_works_count":48,"referenced_works":["https://openalex.org/W1499084133","https://openalex.org/W1556616831","https://openalex.org/W1562077884","https://openalex.org/W1716549302","https://openalex.org/W1763189032","https://openalex.org/W1977802224","https://openalex.org/W1995454352","https://openalex.org/W2000580699","https://openalex.org/W2001390943","https://openalex.org/W2010777779","https://openalex.org/W2011316735","https://openalex.org/W2070809971","https://openalex.org/W2072432549","https://openalex.org/W2114355711","https://openalex.org/W2118638308","https://openalex.org/W2122082385","https://openalex.org/W2148698435","https://openalex.org/W2161444534","https://openalex.org/W2169260566","https://openalex.org/W2345772773","https://openalex.org/W2382988971","https://openalex.org/W2390443808","https://openalex.org/W2415076793","https://openalex.org/W2750970422","https://openalex.org/W2793616504","https://openalex.org/W2972754927","https://openalex.org/W2982666534","https://openalex.org/W2999500215","https://openalex.org/W3012257507","https://openalex.org/W3037558183","https://openalex.org/W3096191613","https://openalex.org/W3155975723","https://openalex.org/W3184862145","https://openalex.org/W4224942167","https://openalex.org/W4226208478","https://openalex.org/W4306156071","https://openalex.org/W4317708817","https://openalex.org/W4382237476","https://openalex.org/W4386226585","https://openalex.org/W4396227366","https://openalex.org/W4396692559","https://openalex.org/W4400520481","https://openalex.org/W4402444249","https://openalex.org/W4403913285","https://openalex.org/W7131099010","https://openalex.org/W7132844030","https://openalex.org/W7147179299","https://openalex.org/W7161051459"],"related_works":[],"abstract_inverted_index":{"Chelicerates":[0],"are":[1],"a":[2,108,112,129,146],"diverse":[3],"group":[4],"of":[5,30,39,67,93,104,135,149,158,186,193,199,202],"terrestrial":[6],"and":[7,13,42,57,78,117,138,141,179,183],"aquatic":[8],"arthropods,":[9],"yet":[10],"their":[11,48],"origin":[12,148],"early":[14,65,97],"evolution":[15,66,201],"remain":[16],"debated1\u20136.":[17],"Among":[18],"different":[19],"hypotheses7\u201311,":[20],"one":[21],"proposes":[22],"that":[23,43,168],"the":[24,34,64,69,90,101,133,156,163,190,200],"front":[25],"appendages,":[26],"known":[27],"as":[28,54,177],"chelicerae,":[29,140],"chelicerates":[31],"evolved":[32],"from":[33,47,72,100,189],"short":[35],"great":[36],"appendages":[37,74,120,144],"(SGAs)":[38],"Cambrian":[40,98],"megacheirans12\u201316":[41],"book":[44,79,150],"gills":[45,80],"originated":[46],"trunk":[49,119,143],"limbs7.":[50],"Although":[51],"taxa":[52],"such":[53,176],"Mollisonia":[55,178],"plenovenatrix7":[56],"Megachelicerax":[58],"cousteaui16":[59],"provide":[60,195],"important":[61],"clues":[62],"to":[63,75,88,173],"chelicerates,":[68],"morphological":[70],"transition":[71],"megacheiran-like":[73],"true":[76,139],"chelicerae":[77],"remains":[81],"unresolved.":[82],"Here":[83],"we":[84],"use":[85],"X-ray":[86],"microtomography":[87],"reveal":[89],"three-dimensional":[91],"anatomy":[92],"Urokodia":[94,106,161,187],"aequalis,":[95],"an":[96,196],"euarthropod":[99],"Chengjiang":[102,191],"biota":[103,192],"China.":[105],"has":[107],"seven-segmented":[109],"head":[110],"with":[111,121,160],"sclerotized":[113],"hypostome,":[114],"pincer-like":[115,126],"SGAs":[116,127,137],"biramous":[118],"overlapping":[122],"exite":[123],"flaps.":[124],"Its":[125],"represent":[128],"bridge":[130],"structure":[131],"between":[132],"appearance":[134],"multisegmented":[136],"its":[142],"support":[145],"megacheiran":[147],"gills.":[151],"Phylogenetic":[152],"analyses":[153,185],"consistently":[154],"confirm":[155],"monophyly":[157],"Cheliceromorpha17,":[159],"being":[162],"earliest-branching":[164],"upper":[165],"stem-group":[166,171],"chelicerate":[167],"links":[169],"lower":[170],"megacheirans":[172],"crownward":[174],"forms":[175],"Megachelicerax.":[180],"Microtomography":[181],"reconstructions":[182],"phylogenetic":[184],"aequalis":[188],"China":[194],"updated":[197],"interpretation":[198],"anatomical":[203],"structures":[204],"in":[205],"chelicerates.":[206]},"counts_by_year":[],"updated_date":"2026-10-15T10:00:46.086208","created_date":"2026-07-02T00:00:00"}