{"id":"https://openalex.org/W7127655428","doi":"https://doi.org/10.3897/zookeys.1268.177040","title":"A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata, Gekkonidae) from Son La Province, Vietnam","display_name":"A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata, Gekkonidae) from Son La Province, Vietnam","publication_year":2026,"publication_date":"2026-02-04","ids":{"openalex":"https://openalex.org/W7127655428","doi":"https://doi.org/10.3897/zookeys.1268.177040","pmid":"https://pubmed.ncbi.nlm.nih.gov/41696482"},"language":"en","primary_location":{"id":"doi:10.3897/zookeys.1268.177040","doi":"https://doi.org/10.3897/zookeys.1268.177040","is_oa":true,"landing_page_url":"https://doi.org/10.3897/zookeys.1268.177040","pdf_url":null,"source":{"id":"https://openalex.org/S199213172","display_name":"ZooKeys","issn_l":"1313-2970","issn":["1313-2970","1313-2989"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","dongbi-d","jufo-1","norway-1","poland-70","russia-white-list-1"],"host_organization":"https://openalex.org/P4310321056","host_organization_name":"Pensoft Publishers","host_organization_lineage":["https://openalex.org/P4310321056"],"host_organization_lineage_names":["Pensoft Publishers"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"ZooKeys","raw_type":"journal-article"},"volume":null,"type":"article","indexed_in":["crossref","doaj","pubmed"],"open_access":{"is_oa":true,"oa_status":"gold","oa_url":"https://doi.org/10.3897/zookeys.1268.177040","any_repository_has_fulltext":true},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5023694807","display_name":"HONG BICH HA","orcid":"https://orcid.org/0000-0003-4777-5463","observed_orcids":["https://orcid.org/0000-0003-4777-5463"]},"institutions":[{"id":"https://openalex.org/I4210123755","display_name":"Vietnam National University of Forestry","ror":"https://ror.org/02jfkxh18","country_code":"VN","type":"education","lineage":["https://openalex.org/I4210123755"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Hong Bich Ha","raw_affiliation_strings":["College of Forestry Biotechnology, Vietnam National University of Forestry, Hanoi, Vietnam"],"raw_orcid":"https://orcid.org/0000-0003-4777-5463","affiliations":[{"raw_affiliation_string":"College of Forestry Biotechnology, Vietnam National University of Forestry, Hanoi, Vietnam","institution_ids":["https://openalex.org/I4210123755"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5109110368","display_name":"Tuoi Thi Hoang","orcid":"https://orcid.org/0000-0002-3564-0601","observed_orcids":["https://orcid.org/0000-0002-3564-0601"]},"institutions":[{"id":"https://openalex.org/I4210123755","display_name":"Vietnam National University of Forestry","ror":"https://ror.org/02jfkxh18","country_code":"VN","type":"education","lineage":["https://openalex.org/I4210123755"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Tuoi Thi Hoang","raw_affiliation_strings":["Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam"],"raw_orcid":"https://orcid.org/0000-0002-3564-0601","affiliations":[{"raw_affiliation_string":"Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam","institution_ids":["https://openalex.org/I4210123755"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5059705201","display_name":"Manh Dac Nguyen","orcid":"https://orcid.org/0009-0002-6075-2274","observed_orcids":["https://orcid.org/0009-0002-6075-2274"]},"institutions":[{"id":"https://openalex.org/I4210123755","display_name":"Vietnam National University of Forestry","ror":"https://ror.org/02jfkxh18","country_code":"VN","type":"education","lineage":["https://openalex.org/I4210123755"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Manh Dac Nguyen","raw_affiliation_strings":["Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam"],"raw_orcid":"https://orcid.org/0009-0002-6075-2274","affiliations":[{"raw_affiliation_string":"Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam","institution_ids":["https://openalex.org/I4210123755"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5137151038","display_name":"Nghia Ha","orcid":"https://orcid.org/0000-0003-2515-8173","observed_orcids":["https://orcid.org/0000-0003-2515-8173"]},"institutions":[],"countries":[],"is_corresponding":false,"raw_author_name":"Nghia Van Ha","raw_affiliation_strings":["Ha Gia Thanh Company Limited, Hanoi, Vietnam"],"raw_orcid":"https://orcid.org/0000-0003-2515-8173","affiliations":[{"raw_affiliation_string":"Ha Gia Thanh Company Limited, Hanoi, Vietnam","institution_ids":[]}]},{"author_position":"last","author":{"id":"https://openalex.org/A5052861262","display_name":"Vinh Quang Luu","orcid":"https://orcid.org/0000-0002-0634-1338","observed_orcids":["https://orcid.org/0000-0002-0634-1338"]},"institutions":[{"id":"https://openalex.org/I4210123755","display_name":"Vietnam National University of Forestry","ror":"https://ror.org/02jfkxh18","country_code":"VN","type":"education","lineage":["https://openalex.org/I4210123755"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Vinh Quang Luu","raw_affiliation_strings":["Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam"],"raw_orcid":"https://orcid.org/0000-0002-0634-1338","affiliations":[{"raw_affiliation_string":"Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Hanoi, Vietnam","institution_ids":["https://openalex.org/I4210123755"]}]}],"institutions":[],"countries_distinct_count":1,"institutions_distinct_count":1,"corresponding_author_ids":[],"corresponding_institution_ids":[],"apc_list":{"value":700,"currency":"EUR","value_usd":754},"apc_paid":{"value":700,"currency":"EUR","value_usd":754},"fwci":0.0,"has_fulltext":false,"cited_by_count":0,"citation_normalized_percentile":{"value":0.06031806,"is_in_top_1_percent":false,"is_in_top_10_percent":false},"cited_by_percentile_year":null,"biblio":{"volume":"1268","issue":null,"first_page":"163","last_page":"188"},"is_retracted":false,"is_paratext":false,"is_xpac":false,"primary_topic":{"id":"https://openalex.org/T10332","display_name":"Amphibian and Reptile Biology","score":0.9829000234603882,"subfield":{"id":"https://openalex.org/subfields/2306","display_name":"Global and Planetary Change"},"field":{"id":"https://openalex.org/fields/23","display_name":"Environmental Science"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},"topics":[{"id":"https://openalex.org/T10332","display_name":"Amphibian and Reptile Biology","score":0.9829000234603882,"subfield":{"id":"https://openalex.org/subfields/2306","display_name":"Global and Planetary Change"},"field":{"id":"https://openalex.org/fields/23","display_name":"Environmental Science"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},{"id":"https://openalex.org/T12319","display_name":"Fish Biology and Ecology Studies","score":0.007899999618530273,"subfield":{"id":"https://openalex.org/subfields/1104","display_name":"Aquatic Science"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}},{"id":"https://openalex.org/T14399","display_name":"Invertebrate Taxonomy and Ecology","score":0.0020000000949949026,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[{"id":"https://openalex.org/keywords/new-species","display_name":"new species","score":0.9470000267028809},{"id":"https://openalex.org/keywords/geckos","display_name":"geckos","score":0.9539999961853027},{"id":"https://openalex.org/keywords/nd2-gene","display_name":"ND2 gene","score":0.7360000014305115},{"id":"https://openalex.org/keywords/morphometrics","display_name":"morphometrics","score":0.4580000042915344},{"id":"https://openalex.org/keywords/meristics","display_name":"meristic characters","score":0.7990000247955322},{"id":"https://openalex.org/keywords/scale-characters","display_name":"scale characters","score":0.5649999976158142},{"id":"https://openalex.org/keywords/karst-habitat","display_name":"karst habitat","score":0.7089999914169312},{"id":"https://openalex.org/keywords/son-la-province","display_name":"Son La province","score":0.8349999785423279},{"id":"https://openalex.org/keywords/species-delimitation","display_name":"species delimitation","score":0.6100000143051147}],"concepts":[{"id":"https://openalex.org/C44465124","wikidata":"https://www.wikidata.org/wiki/Q713623","display_name":"Clade","level":4,"score":0.8582000136375427},{"id":"https://openalex.org/C157369684","wikidata":"https://www.wikidata.org/wiki/Q34740","display_name":"Genus","level":2,"score":0.4853000044822693},{"id":"https://openalex.org/C205649164","wikidata":"https://www.wikidata.org/wiki/Q1071","display_name":"Geography","level":0,"score":0.474700003862381},{"id":"https://openalex.org/C90856448","wikidata":"https://www.wikidata.org/wiki/Q431","display_name":"Zoology","level":1,"score":0.45719999074935913},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.43549999594688416},{"id":"https://openalex.org/C58642233","wikidata":"https://www.wikidata.org/wiki/Q8269924","display_name":"Taxonomy (biology)","level":2,"score":0.41999998688697815},{"id":"https://openalex.org/C78458016","wikidata":"https://www.wikidata.org/wiki/Q840400","display_name":"Evolutionary biology","level":1,"score":0.3621000051498413},{"id":"https://openalex.org/C71640776","wikidata":"https://www.wikidata.org/wiki/Q16521","display_name":"Taxon","level":2,"score":0.35519999265670776},{"id":"https://openalex.org/C182348080","wikidata":"https://www.wikidata.org/wiki/Q16817","display_name":"Karst","level":2,"score":0.3546000123023987},{"id":"https://openalex.org/C140530291","wikidata":"https://www.wikidata.org/wiki/Q1225063","display_name":"Dorsum","level":2,"score":0.33629998564720154},{"id":"https://openalex.org/C193252679","wikidata":"https://www.wikidata.org/wiki/Q242125","display_name":"Phylogenetic tree","level":3,"score":0.31060001254081726},{"id":"https://openalex.org/C90132467","wikidata":"https://www.wikidata.org/wiki/Q171184","display_name":"Phylogenetics","level":3,"score":0.30959999561309814},{"id":"https://openalex.org/C2780861071","wikidata":"https://www.wikidata.org/wiki/Q1062934","display_name":"Character (mathematics)","level":2,"score":0.30379998683929443},{"id":"https://openalex.org/C5274069","wikidata":"https://www.wikidata.org/wiki/Q2285707","display_name":"Categorical variable","level":2,"score":0.2906999886035919},{"id":"https://openalex.org/C2779881321","wikidata":"https://www.wikidata.org/wiki/Q82714","display_name":"Chin","level":2,"score":0.2784999907016754},{"id":"https://openalex.org/C141732470","wikidata":"https://www.wikidata.org/wiki/Q1154642","display_name":"DNA barcoding","level":2,"score":0.2777999937534332},{"id":"https://openalex.org/C18903297","wikidata":"https://www.wikidata.org/wiki/Q7150","display_name":"Ecology","level":1,"score":0.26750001311302185}],"mesh":[],"locations_count":7,"locations":[{"id":"doi:10.3897/zookeys.1268.177040","doi":"https://doi.org/10.3897/zookeys.1268.177040","is_oa":true,"landing_page_url":"https://doi.org/10.3897/zookeys.1268.177040","pdf_url":null,"source":{"id":"https://openalex.org/S199213172","display_name":"ZooKeys","issn_l":"1313-2970","issn":["1313-2970","1313-2989"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","dongbi-d","jufo-1","norway-1","poland-70","russia-white-list-1"],"host_organization":"https://openalex.org/P4310321056","host_organization_name":"Pensoft Publishers","host_organization_lineage":["https://openalex.org/P4310321056"],"host_organization_lineage_names":["Pensoft Publishers"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"ZooKeys","raw_type":"journal-article"},{"id":"pmid:41696482","doi":null,"is_oa":false,"landing_page_url":"https://pubmed.ncbi.nlm.nih.gov/41696482","pdf_url":null,"source":{"id":"https://openalex.org/S4306525036","display_name":"PubMed","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I1299303238","host_organization_name":"National Institutes of Health","host_organization_lineage":["https://openalex.org/I1299303238"],"host_organization_lineage_names":[],"type":"repository"},"license":null,"license_id":null,"version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"ZooKeys","raw_type":"Journal Article"},{"id":"pmh:oai:europepmc.org:11702443","doi":null,"is_oa":true,"landing_page_url":"https://www.ncbi.nlm.nih.gov/pmc/articles/12895182","pdf_url":null,"source":{"id":"https://openalex.org/S4306400806","display_name":"Europe PMC (PubMed Central)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I1303153112","host_organization_name":"European Bioinformatics Institute","host_organization_lineage":["https://openalex.org/I1303153112"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Text"},{"id":"pmh:10.3897/zookeys.1268.177040","doi":null,"is_oa":true,"landing_page_url":"https://doi.org/10.3897/zookeys.1268.177040","pdf_url":null,"source":{"id":"https://openalex.org/S199213172","display_name":"ZooKeys","issn_l":"1313-2970","issn":["1313-2970","1313-2989"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","dongbi-d","jufo-1","norway-1","poland-70","russia-white-list-1"],"host_organization":"https://openalex.org/P4310321056","host_organization_name":"Pensoft Publishers","host_organization_lineage":["https://openalex.org/P4310321056"],"host_organization_lineage_names":["Pensoft Publishers"],"type":"journal"},"license":"other-oa","license_id":"https://openalex.org/licenses/other-oa","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"ZooKeys 1268: 163-188","raw_type":"Research Article"},{"id":"pmh:oai:biodiversitylibrary.org:part/442134","doi":null,"is_oa":true,"landing_page_url":"https://www.biodiversitylibrary.org/partpdf/442134","pdf_url":null,"source":{"id":"https://openalex.org/S4306402618","display_name":"Biodiversity Heritage Library (Smithsonian Institution)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I103187081","host_organization_name":"Smithsonian Institution","host_organization_lineage":["https://openalex.org/I103187081"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":null,"raw_type":"Article"},{"id":"pmh:oai:doaj.org/article:72e822b1277b4919939be40df66d3416","doi":null,"is_oa":true,"landing_page_url":"https://doaj.org/article/72e822b1277b4919939be40df66d3416","pdf_url":null,"source":{"id":"https://openalex.org/S4306401280","display_name":"DOAJ (DOAJ: Directory of Open Access Journals)","issn_l":null,"issn":null,"is_oa":false,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":null,"host_organization_name":null,"host_organization_lineage":[],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by-sa","license_id":"https://openalex.org/licenses/cc-by-sa","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"ZooKeys, Vol 1268, Iss , Pp 163-188 (2026)","raw_type":"article"},{"id":"pmh:oai:pubmedcentral.nih.gov:12895182","doi":null,"is_oa":true,"landing_page_url":"https://pmc.ncbi.nlm.nih.gov/articles/PMC12895182/","pdf_url":null,"source":{"id":"https://openalex.org/S2764455111","display_name":"PubMed Central","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I1299303238","host_organization_name":"National Institutes of Health","host_organization_lineage":["https://openalex.org/I1299303238"],"host_organization_lineage_names":[],"type":"repository"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"Zookeys","raw_type":"Text"}],"best_oa_location":{"id":"doi:10.3897/zookeys.1268.177040","doi":"https://doi.org/10.3897/zookeys.1268.177040","is_oa":true,"landing_page_url":"https://doi.org/10.3897/zookeys.1268.177040","pdf_url":null,"source":{"id":"https://openalex.org/S199213172","display_name":"ZooKeys","issn_l":"1313-2970","issn":["1313-2970","1313-2989"],"is_oa":true,"is_in_doaj":true,"is_core":true,"listed_in":["cwts-core","doaj","dongbi-d","jufo-1","norway-1","poland-70","russia-white-list-1"],"host_organization":"https://openalex.org/P4310321056","host_organization_name":"Pensoft Publishers","host_organization_lineage":["https://openalex.org/P4310321056"],"host_organization_lineage_names":["Pensoft Publishers"],"type":"journal"},"license":"cc-by","license_id":"https://openalex.org/licenses/cc-by","version":"publishedVersion","is_accepted":true,"is_published":true,"raw_source_name":"ZooKeys","raw_type":"journal-article"},"sustainable_development_goals":[],"sustainable_development_goals_aurora":[{"score":0.5463253259658813,"id":"https://metadata.un.org/sdg/15","display_name":"Life on land"}],"study_designs":[],"awards":[],"funders":[],"has_content":{"pdf":false,"grobid_xml":false},"content_urls":null,"referenced_works_count":39,"referenced_works":["https://openalex.org/W381842185","https://openalex.org/W1487144146","https://openalex.org/W1977601452","https://openalex.org/W1983102293","https://openalex.org/W1996655166","https://openalex.org/W2014250557","https://openalex.org/W2027332124","https://openalex.org/W2028302194","https://openalex.org/W2029089143","https://openalex.org/W2031211350","https://openalex.org/W2078231496","https://openalex.org/W2089874952","https://openalex.org/W2111647009","https://openalex.org/W2141913814","https://openalex.org/W2148698435","https://openalex.org/W2278234919","https://openalex.org/W2336400828","https://openalex.org/W2614081736","https://openalex.org/W2751251894","https://openalex.org/W2802817419","https://openalex.org/W2890973362","https://openalex.org/W2951464304","https://openalex.org/W3013091970","https://openalex.org/W3036875658","https://openalex.org/W3047359488","https://openalex.org/W3167208384","https://openalex.org/W4223649474","https://openalex.org/W4291399178","https://openalex.org/W4381611075","https://openalex.org/W4381832324","https://openalex.org/W4399538195","https://openalex.org/W4400900620","https://openalex.org/W4403603768","https://openalex.org/W4404233009","https://openalex.org/W4405670661","https://openalex.org/W4406314525","https://openalex.org/W4410278754","https://openalex.org/W4412941235","https://openalex.org/W4413722879"],"related_works":[],"abstract_inverted_index":{"An":[0],"integrative":[1],"analysis":[2,92],"revealed":[3],"a":[4,62],"new":[5,126],"species":[6,115,128,144],"of":[7,44,67,124,133,143,145],"the":[8,45,68,117,130,134,141,150],"Hemiphyllodactylus":[9,34,127],"typus":[10,46],"group":[11],"collected":[12],"from":[13,56,75,112,137],"local":[14],"house":[15],"walls":[16],"and":[17,86,97,122,139],"surrounding":[18],"karst":[19],"formations":[20],"in":[21,106,116,149],"Pa":[22],"Kha":[23],"1":[24],"Village,":[25],"Long":[26],"Phieng":[27],"Commune,":[28],"Son":[29],"La":[30],"Province,":[31],"northwestern":[32],"Vietnam.":[33],"pakhaensis":[35],"sp.":[36],"nov":[37],".":[38],"is":[39],"recovered":[40,101],"within":[41],"Clade":[42,118,135],"4":[43,136],"group,":[47],"exhibiting":[48],"an":[49],"uncorrected":[50],"pairwise":[51],"distance":[52],"greater":[53],"than":[54],"6.65%":[55],"all":[57],"other":[58],"congeners":[59,77],"based":[60],"on":[61],"1,043":[63],"base":[64],"pair":[65],"segment":[66],"ND2":[69],"gene.":[70],"It":[71],"can":[72],"be":[73],"distinguished":[74],"its":[76,102],"by":[78],"body":[79],"size,":[80],"chin":[81],"scales,":[82,84],"circumnasal":[83],"dorsal":[85],"ventral":[87],"scales.":[88],"A":[89],"multiple":[90],"factor":[91],"using":[93],"normalized":[94],"morphometric,":[95],"meristic,":[96],"categorical":[98],"character":[99],"types":[100],"unique,":[103],"non\u2013overlapping":[104],"placement":[105],"morphospace":[107],"as":[108],"statistically":[109],"significantly":[110],"different":[111],"closely":[113],"related":[114],"4.":[119],"The":[120],"discovery":[121],"description":[123],"this":[125,146],"represent":[129],"first":[131],"record":[132],"Vietnam":[138],"increase":[140],"number":[142],"genus":[147],"recorded":[148],"country":[151],"to":[152],"12.":[153]},"counts_by_year":[],"updated_date":"2026-10-14T16:39:01.820549","created_date":"2026-02-06T00:00:00"}