{"id":"https://openalex.org/W6968201848","doi":"https://doi.org/10.5281/zenodo.15376293","title":"Anotogaster klossi Fraser 1919","display_name":"Anotogaster klossi Fraser 1919","publication_year":2025,"publication_date":"2025-05-07","ids":{"openalex":"https://openalex.org/W6968201848","doi":"https://doi.org/10.5281/zenodo.15376293"},"language":"en","primary_location":{"id":"pmh:oai:zenodo.org:15376294","doi":null,"is_oa":true,"landing_page_url":"https://doi.org/10.5281/zenodo.15376294","pdf_url":null,"source":{"id":"https://openalex.org/S4306400562","display_name":"Zenodo (CERN European Organization for Nuclear Research)","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I67311998","host_organization_name":"European Organization for Nuclear Research","host_organization_lineage":["https://openalex.org/I67311998"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"Description of the larva of Anotogaster klossi Fraser, 1919 (Odonata, Cordulegastroidea, Cordulegastridae), with a note on the known larvae of the genus Anotogaster Selys, 1854, pp. 349-363 in Zootaxa, 5632(2), 350-356, (2025-05-07)","raw_type":"info:eu-repo/semantics/other"},"volume":null,"type":"other","indexed_in":["datacite"],"open_access":{"is_oa":true,"oa_status":"green","oa_url":"https://doi.org/10.5281/zenodo.15376294","any_repository_has_fulltext":true},"authorships":[{"author_position":"first","author":{"id":"https://openalex.org/A5036732431","display_name":"Tosaphol Saetung Keetapithchayakul","orcid":"https://orcid.org/0000-0001-7565-4701","observed_orcids":["https://orcid.org/0000-0001-7565-4701"]},"institutions":[{"id":"https://openalex.org/I170238339","display_name":"Duy Tan University","ror":"https://ror.org/05ezss144","country_code":"VN","type":"education","lineage":["https://openalex.org/I170238339"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Keetapithchayakul, Tosaphol Saetung","raw_affiliation_strings":["The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam","institution_ids":["https://openalex.org/I170238339"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5152983075","display_name":"Junggon Kim","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I170238339","display_name":"Duy Tan University","ror":"https://ror.org/05ezss144","country_code":"VN","type":"education","lineage":["https://openalex.org/I170238339"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Kim, Junggon","raw_affiliation_strings":["The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam","institution_ids":["https://openalex.org/I170238339"]}]},{"author_position":"middle","author":{"id":"https://openalex.org/A5146858836","display_name":"Nguyen Ba Vu Lam","orcid":null,"observed_orcids":[]},"institutions":[],"countries":[],"is_corresponding":false,"raw_author_name":"Lam, Nguyen Ba Vu","raw_affiliation_strings":["Science and International Cooperation Division, Bach Ma National Park, Phu Loc, Hue, Vietnam"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"Science and International Cooperation Division, Bach Ma National Park, Phu Loc, Hue, Vietnam","institution_ids":[]}]},{"author_position":"last","author":{"id":null,"display_name":"Phan, Quoc Toan","orcid":null,"observed_orcids":[]},"institutions":[{"id":"https://openalex.org/I170238339","display_name":"Duy Tan University","ror":"https://ror.org/05ezss144","country_code":"VN","type":"education","lineage":["https://openalex.org/I170238339"]}],"countries":["VN"],"is_corresponding":false,"raw_author_name":"Phan, Quoc Toan","raw_affiliation_strings":["The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam"],"raw_orcid":null,"affiliations":[{"raw_affiliation_string":"The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam","institution_ids":["https://openalex.org/I170238339"]}]}],"institutions":[],"countries_distinct_count":1,"institutions_distinct_count":1,"corresponding_author_ids":[],"corresponding_institution_ids":[],"apc_list":null,"apc_paid":null,"fwci":null,"has_fulltext":false,"cited_by_count":0,"citation_normalized_percentile":null,"cited_by_percentile_year":null,"biblio":{"volume":null,"issue":null,"first_page":null,"last_page":null},"is_retracted":false,"is_paratext":false,"is_xpac":true,"primary_topic":{"id":"https://openalex.org/T10669","display_name":"Freshwater macroinvertebrate diversity and ecology","score":0.36820000410079956,"subfield":{"id":"https://openalex.org/subfields/2303","display_name":"Ecology"},"field":{"id":"https://openalex.org/fields/23","display_name":"Environmental Science"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},"topics":[{"id":"https://openalex.org/T10669","display_name":"Freshwater macroinvertebrate diversity and ecology","score":0.36820000410079956,"subfield":{"id":"https://openalex.org/subfields/2303","display_name":"Ecology"},"field":{"id":"https://openalex.org/fields/23","display_name":"Environmental Science"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},{"id":"https://openalex.org/T13578","display_name":"Stonefly species taxonomy and ecology","score":0.13950000703334808,"subfield":{"id":"https://openalex.org/subfields/2212","display_name":"Ocean Engineering"},"field":{"id":"https://openalex.org/fields/22","display_name":"Engineering"},"domain":{"id":"https://openalex.org/domains/3","display_name":"Physical Sciences"}},{"id":"https://openalex.org/T12329","display_name":"Hemiptera Insect Studies","score":0.11400000005960464,"subfield":{"id":"https://openalex.org/subfields/1105","display_name":"Ecology, Evolution, Behavior and Systematics"},"field":{"id":"https://openalex.org/fields/11","display_name":"Agricultural and Biological Sciences"},"domain":{"id":"https://openalex.org/domains/1","display_name":"Life Sciences"}}],"keywords":[{"id":"https://openalex.org/keywords/larval-morphology","display_name":"larval morphology","score":0.5019999742507935},{"id":"https://openalex.org/keywords/larval-description","display_name":"larval description","score":0.8529999852180481},{"id":"https://openalex.org/keywords/larval-exuviae","display_name":"larval exuviae","score":0.6669999957084656},{"id":"https://openalex.org/keywords/larval-instars","display_name":"larval instars","score":0.6119999885559082},{"id":"https://openalex.org/keywords/larval-pigmentation","display_name":"larval coloration","score":0.5109999775886536},{"id":"https://openalex.org/keywords/larval-setae","display_name":"larval setae","score":0.5329999923706055},{"id":"https://openalex.org/keywords/larval-mouthparts","display_name":"larval mouthparts","score":0.5139999985694885}],"concepts":[{"id":"https://openalex.org/C158710501","wikidata":"https://www.wikidata.org/wiki/Q126780","display_name":"Seta","level":3,"score":0.810699999332428},{"id":"https://openalex.org/C105702510","wikidata":"https://www.wikidata.org/wiki/Q514","display_name":"Anatomy","level":1,"score":0.6639000177383423},{"id":"https://openalex.org/C140530291","wikidata":"https://www.wikidata.org/wiki/Q1225063","display_name":"Dorsum","level":2,"score":0.5160999894142151},{"id":"https://openalex.org/C2775889985","wikidata":"https://www.wikidata.org/wiki/Q360768","display_name":"Labrum","level":3,"score":0.49709999561309814},{"id":"https://openalex.org/C3020737034","wikidata":"https://www.wikidata.org/wiki/Q4771399","display_name":"Anterior surface","level":2,"score":0.453000009059906},{"id":"https://openalex.org/C127313418","wikidata":"https://www.wikidata.org/wiki/Q1069","display_name":"Geology","level":0,"score":0.4489000141620636},{"id":"https://openalex.org/C37054046","wikidata":"https://www.wikidata.org/wiki/Q641888","display_name":"Elevation (ballistics)","level":2,"score":0.3950999975204468},{"id":"https://openalex.org/C2777586403","wikidata":"https://www.wikidata.org/wiki/Q4243098","display_name":"Hook","level":2,"score":0.3889000117778778},{"id":"https://openalex.org/C2778539849","wikidata":"https://www.wikidata.org/wiki/Q7596467","display_name":"Stadium","level":2,"score":0.35100001096725464},{"id":"https://openalex.org/C2778275279","wikidata":"https://www.wikidata.org/wiki/Q158717","display_name":"Lycaenidae","level":3,"score":0.3499999940395355},{"id":"https://openalex.org/C86803240","wikidata":"https://www.wikidata.org/wiki/Q420","display_name":"Biology","level":0,"score":0.34360000491142273},{"id":"https://openalex.org/C114138010","wikidata":"https://www.wikidata.org/wiki/Q7684542","display_name":"Tapering","level":2,"score":0.3172000050544739},{"id":"https://openalex.org/C2778090171","wikidata":"https://www.wikidata.org/wiki/Q250234","display_name":"Baetidae","level":3,"score":0.31679999828338623},{"id":"https://openalex.org/C48880136","wikidata":"https://www.wikidata.org/wiki/Q864673","display_name":"Crest","level":2,"score":0.30250000953674316},{"id":"https://openalex.org/C41451933","wikidata":"https://www.wikidata.org/wiki/Q13415392","display_name":"Appendage","level":2,"score":0.28870001435279846},{"id":"https://openalex.org/C2780993040","wikidata":"https://www.wikidata.org/wiki/Q46169","display_name":"National park","level":2,"score":0.2606000006198883},{"id":"https://openalex.org/C3020541326","wikidata":"https://www.wikidata.org/wiki/Q1796666","display_name":"Body surface","level":2,"score":0.2572000026702881},{"id":"https://openalex.org/C186348155","wikidata":"https://www.wikidata.org/wiki/Q7975538","display_name":"Waves and shallow water","level":2,"score":0.25690001249313354}],"mesh":[],"locations_count":3,"locations":[{"id":"pmh:oai:zenodo.org:15376294","doi":null,"is_oa":true,"landing_page_url":"https://doi.org/10.5281/zenodo.15376294","pdf_url":null,"source":{"id":"https://openalex.org/S4306400562","display_name":"Zenodo (CERN European Organization for Nuclear Research)","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I67311998","host_organization_name":"European Organization for Nuclear Research","host_organization_lineage":["https://openalex.org/I67311998"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"Description of the larva of Anotogaster klossi Fraser, 1919 (Odonata, Cordulegastroidea, Cordulegastridae), with a note on the known larvae of the genus Anotogaster Selys, 1854, pp. 349-363 in Zootaxa, 5632(2), 350-356, (2025-05-07)","raw_type":"info:eu-repo/semantics/other"},{"id":"doi:10.5281/zenodo.15376293","doi":"https://doi.org/10.5281/zenodo.15376293","is_oa":true,"landing_page_url":"https://doi.org/10.5281/zenodo.15376293","pdf_url":null,"source":{"id":"https://openalex.org/S4306400562","display_name":"Zenodo (CERN European Organization for Nuclear Research)","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I67311998","host_organization_name":"European Organization for Nuclear Research","host_organization_lineage":["https://openalex.org/I67311998"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":null,"is_accepted":false,"is_published":null,"raw_source_name":null,"raw_type":"article-journal"},{"id":"doi:10.5281/zenodo.15376294","doi":"https://doi.org/10.5281/zenodo.15376294","is_oa":true,"landing_page_url":"https://doi.org/10.5281/zenodo.15376294","pdf_url":null,"source":{"id":"https://openalex.org/S4306400562","display_name":"Zenodo (CERN European Organization for Nuclear Research)","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I67311998","host_organization_name":"European Organization for Nuclear Research","host_organization_lineage":["https://openalex.org/I67311998"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":null,"is_accepted":false,"is_published":null,"raw_source_name":null,"raw_type":"article-journal"}],"best_oa_location":{"id":"pmh:oai:zenodo.org:15376294","doi":null,"is_oa":true,"landing_page_url":"https://doi.org/10.5281/zenodo.15376294","pdf_url":null,"source":{"id":"https://openalex.org/S4306400562","display_name":"Zenodo (CERN European Organization for Nuclear Research)","issn_l":null,"issn":null,"is_oa":true,"is_in_doaj":false,"is_core":false,"listed_in":[],"host_organization":"https://openalex.org/I67311998","host_organization_name":"European Organization for Nuclear Research","host_organization_lineage":["https://openalex.org/I67311998"],"host_organization_lineage_names":[],"type":"repository"},"license":"public-domain","license_id":"https://openalex.org/licenses/public-domain","version":"submittedVersion","is_accepted":false,"is_published":false,"raw_source_name":"Description of the larva of Anotogaster klossi Fraser, 1919 (Odonata, Cordulegastroidea, Cordulegastridae), with a note on the known larvae of the genus Anotogaster Selys, 1854, pp. 349-363 in Zootaxa, 5632(2), 350-356, (2025-05-07)","raw_type":"info:eu-repo/semantics/other"},"sustainable_development_goals":[],"sustainable_development_goals_aurora":[],"study_designs":[],"awards":[],"funders":[],"has_content":{"grobid_xml":false,"pdf":false},"content_urls":null,"referenced_works_count":0,"referenced_works":[],"related_works":[],"abstract_inverted_index":{"Description":[0,108,1094],"of":[1,3,182,206,225,256,273,357,377,397,413,416,424,436,468,491,521,524,537,558,564,574,581,665,686,697,738,777,783,798,803,836,846,853,860,873,881,887,923,928,939,945,954,963,967,986,992,1011,1035,1070,1095,1125,1136,1148,1199,1309,1328],"larva":[2,1251],"Anotogaster":[4],"klossi":[5,1127],"Fraser,":[6],"1919":[7],"(Figures":[8],"1\u20136,":[9],"7D,":[10,711,902],"8,":[11],"9F,":[12],"10F)":[13],"Material":[14],"examined.":[15],"VIETNAM.":[16],"2":[17,59,61,81,83],"exuviae:":[18],"1":[19,21,111,277,607],"\u2642,":[20],"\u2640":[22,112],"(last":[23,113],"stadium":[24,57,114],"larvae":[25,1124,1164,1192,1198,1273],"when":[26],"collected,":[27],"then":[28],"reared":[29],"in":[30,212,361,429,476,508,970,1215,1253,1266],"laboratory),":[31],"Do":[32,85],"Quyen":[33,86],"waterfall,":[34,87],"Bach":[35,88,117],"Ma":[36,89,118],"National":[37,90,119],"Park":[38,91],"(16\u00b011\u201933.8\u201dN":[39,92],"107\u00b050\u201954.1\u201dE,":[40,93],"elevation":[41,67,94],"1160":[42,95],"m":[43,69,96,654],"a.s.l.),":[44,70,97],"Thua":[45,98],"Thien-Hue":[46,99],"province,":[47,76,100],"7/IV/2024,":[48,101],"Q.T.":[49,102],"Phan":[50,103,1212,1244],"&":[51,104,1213,1219,1238,1243],"T.S.":[52,78,105],"Keetapithchayakul":[53,79,106,1239,1242],"leg.;":[54],"8":[55],"last":[56],"larvae:":[58],"\u2642\u2642,":[60,82],"\u2640\u2640,":[62,84],"Tr\u2019":[63],"Hy":[64],"(15\u00b049\u201911.0\u201dN":[65],"107\u00b021\u201945.9\u201dE,":[66],"1261":[68],"Tay":[71],"Giang":[72],"District,":[73],"Quang":[74],"Nam":[75],"5/VIII/2024,":[77],"leg.,":[80],"leg.":[107],"(based":[109],"on":[110,175,241,260,328,336,348,364,374,448,483,494,504,570,598,704,726,755,770,892,919,995,1061,1081],"larvae)":[115],"from":[116],"Park)":[120],"Large":[121],"size":[122,430,560,1104],"(48\u201351":[123],"mm":[124,1106],"total":[125,1107],"length),":[126,1108],"female":[127],"significantly":[128],"larger":[129,876],"than":[130,157,714,822,877],"male,":[131],"body":[132],"robust,":[133,840,854,888],"setose":[134,593],"and":[135,146,178,198,210,254,290,296,314,326,420,432,444,458,487,561,603,609,791,800,829,848,857,895,914,961,1000,1015,1044,1063,1121,1158,1171,1232,1263,1305,1339],"tapering":[136,906],"caudally,":[137],"with":[138,172,222,231,237,250,293,298,317,323,333,355,367,371,389,410,439,455,478,578,606,623,630,642,650,680,700,723,751,767,773,780,816,826,838,850,868,884,910,925,932,948,957,981,989,997,1022,1057,1067,1077,1114,1132,1185,1196],"dense":[139,180,489,579,724,768,817,915,1078],"yellow":[140,148],"setae,":[141],"caudal":[142,1335],"appendages":[143,1039],"sharply":[144,617],"pointed,":[145,618,1046,1087],"coloration":[147,1176],"orange":[149],"to":[150,465,587,1182,1323],"reddish":[151],"brown":[152],"(Fig.":[153,166,169,194,215,248,303,311,341,352,400,453,471,529,546,621,677,807,975,1027,1040,1139,1270,1342],"1).":[154,167],"Head.":[155],"Wider":[156],"long,":[158,226,318,368,752,916],"about":[159],"2.2":[160],"times":[161],"as":[162,164,539,743,1053,1055,1089,1091,1178],"wide":[163],"long":[165,183,232,258,324,358,372,391,446,457,582,611,652,674,818,933,950,959,1043,1054,1059,1079,1090],"Labrum":[168],"2C,":[170,195],"F),":[171],"short":[173,460,608,855,993,998],"SS":[174,184,259,325,359,373,447,725,754,769,828,918,930,960,1060,1080],"external":[176],"surface,":[177,1076],"a":[179,223,362,440,479,488,509,540,567,660,744,781,851,858,885,926,971,990,1019,1068,1115,1267,1275],"brush":[181],"along":[185,1074],"the":[186,268,484,498,544,548,688,705,1197,1254,1321,1325,1329,1334],"widely":[187],"concave":[188,475],"ventral":[189,375,386,417,449,584,695],"margin,":[190,330,486],"flattened":[191],"ventrally.":[192],"Clypeus":[193],"F)":[196,531],"glabrous":[197,209,604],"moderately":[199,219,812],"striated.":[200],"In":[201],"frontal":[202],"view,":[203],"anterior":[204,217,535,729,790],"surface":[205,235,376,387,585],"frons,":[207],"flat,":[208],"striated;":[211],"dorsal":[213,234,337,425,572,599,684,775,1118,1264],"view":[214],"2D),":[216],"margin":[218,550,573,664,730,740,759,797,894,938,1034],"convex,":[220],"beset":[221,750],"row":[224,356,418,426,490,852,859,886,927,962,991,1069],"stout,":[227,319],"thick":[228,320],"PS,":[229,295,321,994],"intermingled":[230,322,370,931],"SS,":[233,913,934],"covered":[236,332,388,716,815,909,979],"small":[238,251,492,591,698,912,929,982,1129,1151,1168],"scale-like":[239,252,334,720],"setae":[240,253,335,452,462],"distal":[242,832,1326],"half,":[243],"basal":[244,801],"half":[245,802],"bare.":[246],"Vertex":[247],"2D)":[249],"tuff":[255],"very":[257],"both":[261,1303],"sides":[262],"among":[263],"bordering":[264],"ocelli.":[265],"Antennae":[266],"7-segmented,":[267],"A3":[269],"longest,":[270],"relative":[271],"length":[272],"antennomeres":[274],"0.55:":[275],"0.61:":[276,282],"(1.09":[278],"mm):":[279],"0.53:":[280,281],"0.40,":[283],"(Figs":[284,381,709,899,1188,1311],"2A,":[285,304],"C,":[286],"D,":[287],"3A),":[288],"A1":[289],"A2":[291],"barrel-shaped,":[292],"scattered":[294,299,445,827,896,935],"A1\u20137":[297],"SS.":[300,675,897],"Compound":[301],"eyes":[302],"C)":[305,384,402],"poorly":[306,343],"developed,":[307,344,533],"protruding":[308],"latero-dorsally.":[309],"Occiput":[310],"2A)":[312,342],"large":[313,784,1103],"laterally":[315],"rounded,":[316],"SPS":[327,830,856,891,1073],"lateral":[329,408,952,965,1075],"densely":[331,717],"surface;":[338,450],"postocular":[339],"lobe":[340],"forming":[345,553,659,760,1018],"slightly":[346,406,474,824,875,941,1049,1331],"straight":[347],"posterior":[349,739,758,764,792,796,937,984,1033],"margin.":[350],"Genae":[351],"2B,":[353],"E)":[354],"arranged":[360],"line":[363],"inner":[365,1082],"margin;":[366],"SPS,":[369,414,583],"compound":[378],"eyes.":[379],"Labium":[380],"1B,":[382],"3B,":[383,401],"spoon-shaped,":[385],"delicate,":[390],"SS;":[392,819,861,951],"prementum-postmentum":[393],"articulation":[394],"reaching":[395,795],"middle":[396,736],"metacoxae.":[398],"Prementum":[399],"subpentagonal,":[403],"latero-basal":[404],"margins":[405,409,506,734,922,966,985],"concave,":[407],"two":[411,588],"rows":[412,580],"those":[415,423],"minute":[419,719],"same":[421,689],"size,":[422,690,699],"small,":[427,513,656],"increasing":[428],"apically":[431],"meeting":[433],"at":[434,519],"base":[435,520],"palp\u2019s":[437],"articulation;":[438],"longitudinal,":[441],"central":[442],"sulcus":[443],"premental":[451,461],"3B)":[454],"7":[456,681],"3\u20134":[459],"(7+3,":[463],"7+4)":[464],"each":[466,522,575],"side":[467],"midline;":[469],"ligula":[470],"3D)":[472],"poorly-developed,":[473],"shape,":[477,1048],"V-shaped":[480],"median":[481,499,525],"process":[482,500,518],"apical":[485,495,549,691,706],"PS":[493,999],"border":[496],"excepting":[497],"which":[501,565],"is":[502],"serrulated":[503],"mesal":[505,893],"ending":[507,969],"sharp":[510],"tip;":[511],"additional":[512],"serrulated,":[514],"blunted":[515,842,890],"or":[516,1134,1259,1284,1291],"pointed":[517,556,1340],"branch":[523],"process;":[526],"labial":[527],"palp":[528,576],"3E,":[530],"strongly":[532,905],"covering":[534,543,601],"part":[536,737],"head":[538],"mask,":[541],"not":[542],"frons":[545],"8B),":[547],"irregularly":[551],"toothed,":[552],"11\u201314":[554],"short,":[555,841,889],"hooks":[557],"different":[559],"robustness,":[562],"all":[563,882],"have":[566],"fine":[568,917],"serration":[569],"margins;":[571,1083],"fringed":[577],"dividing":[586],"regions":[589],"by":[590,718,1281,1302,1320],"setae:":[592],"region":[594],"(scattered":[595],"stout":[596,1072],"SS)":[597],"side,":[600],"3/4":[602],"region;":[605],"5":[610],"palpal":[612],"setae;":[613],"movable":[614],"hook":[615],"slender,":[616],"smooth.":[619],"Mandibles":[620],"4A\u2013H)":[622],"formula:":[624,863],"L":[625],"12340":[626],"a(m":[627,639],"1,2,3)":[628],"b,":[629,641],"incisors":[631,643],"1>":[632,644],"2>":[633,645],"3>":[634,646],"4;":[635],"R":[636],"12345":[637],"y":[638],"1,2,3,4,5)":[640],"4>5,":[647],"asymmetrical,":[648],"robust":[649],"well-developed":[651,869],"teeth,":[653,682],"denticles":[655],"blunt,":[657],"semi-fused":[658],"continuous":[661,1020],"ridge;":[662],"outer":[663,1062],"mandible":[666],"smoothly":[667],"angulate":[668],"without":[669],"obvious":[670],"spines,":[671],"bearing":[672],"antero-laterally":[673],"Maxilla":[676],"4I\u2013K)":[678],"galeolacinia":[679],"4":[683],"teeth":[685,696],"approximately":[687],"tooth":[692,703],"largest,":[693],"3":[694],"one":[701,1013],"smallest":[702],"tooth.":[707],"Thorax":[708],"1,":[710,900],"8).":[712,903],"Narrower":[713],"head,":[715],"setae.":[721],"Prothorax":[722],"propleura,":[727],"pronotum":[728,1262],"almost":[731],"straight,":[732],"posterolateral":[733,756],"wavy,":[735],"produced":[741],"caudally":[742],"\u201cpull":[745],"flap\u201d,":[746],"transverse":[747,785],"pronotal":[748],"disc":[749],"stiff":[753],"margins,":[757],"subtle":[761],"ridge":[762],"across":[763],"border.":[765],"Pterothorax":[766],"pleura;":[771],"notum":[772],"white":[774],"area":[776,944,953],"mesopleura":[778,1265],"shield-like,":[779],"pair":[782],"spiracles.":[786],"Wing":[787],"sheaths":[788,794,1258],"divergent;":[789],"wing":[793,1257],"S3":[799],"S4,":[804],"respectively.":[805],"Legs":[806],"5)":[808],"coxa":[809],"robust;":[810],"femora":[811],"thick,":[813],"elongated,":[814],"tibia":[820],"slender":[821],"femora,":[823],"curved,":[825],"towards":[831],"end;":[833],"tibial":[834,844],"comb":[835,845],"foreleg":[837],"numerous":[839],"SPS;":[843,936,964,983,1001],"mid-":[847],"hindlegs":[849],"tarsal":[862],"3-3-3,":[864],"pretarsal":[865,871],"claw":[866,872],"simple,":[867],"empodium,":[870],"hindleg":[874],"other":[878],"legs;":[879],"tarsi":[880],"legs":[883],"Abdomen":[898],"6,":[901],"Cylindrical,":[904],"caudally.":[907],"Terga":[908],"abundant,":[911],"sides.":[920],"Posterior":[921],"S2\u20139":[924],"S10":[940],"serrate.":[942],"Lateral":[943],"terga":[946,955],"S1\u20135":[947],"dense,":[949,958],"S5\u201310":[956],"S8\u20139":[968],"convergent,":[972],"acute":[973],"spines":[974],"6D).":[976],"Sterna":[977],"S1\u201310":[978],"abundantly":[980],"sterna":[987,1002],"S1\u20136":[988],"S7\u201310":[996],"2\u20138":[1003],"divided":[1004],"into":[1005,1295],"three":[1006],"sternites,":[1007],"sternum":[1008,1016,1036],"S9":[1009],"consisting":[1010],"only":[1012],"sternite,":[1014],"10":[1017],"ring":[1021],"respective":[1023],"tergum.":[1024],"Female":[1025],"gonapophyses":[1026,1110],"6D\u2013E)":[1028],"well-developed,":[1029],"smoothed,":[1030],"surpassing":[1031],"beyond":[1032],"S9.":[1037],"Caudal":[1038],"6A\u2013C),":[1041],"tips":[1042],"acutely":[1045,1086],"pyramidal":[1047],"downcurved.":[1050],"Epiproct":[1051],"0.94x":[1052],"paraprocts,":[1056],"abundant":[1058],"medial":[1064],"margins.":[1065],"Paraprocts":[1066],"six":[1071],"cerci":[1084],"conical,":[1085],"0.20":[1088],"epiproct":[1092,1113],"length.":[1093],"male":[1096,1109,1112],"larva.":[1097],"As":[1098],"female,":[1099],"unless":[1100],"otherwise":[1101],"stated:":[1102],"(41\u201346":[1105],"poorly-developed;":[1111],"basal,":[1116],"mound-like":[1117],"tubercle.":[1119],"Habitat":[1120],"biology.":[1122],"The":[1123,1141,1163,1174,1191,1272],"A.":[1126],"inhabit":[1128],"forest":[1130],"streams,":[1131],"pools":[1133],"areas":[1135],"slow-moving":[1137],"water":[1138],"7A\u2013C).":[1140],"streambed":[1142],"where":[1143],"collections":[1144],"were":[1145,1193],"made":[1146],"consisted":[1147],"mud/silt":[1149],"(40%),":[1150],"stones/pebbles/":[1152],"gravel/sand":[1153],"(20%),":[1154],"leaf":[1155,1172],"litter":[1156],"(25%),":[1157],"riparian/root":[1159],"tree":[1160],"debris":[1161],"(15%).":[1162],"hide":[1165],"themselves":[1166],"under":[1167],"rocks,":[1169],"slit":[1170],"litter.":[1173],"larval":[1175],"acts":[1177],"camouflage,":[1179],"allowing":[1180],"them":[1181],"blend":[1183],"seamlessly":[1184,1294],"their":[1186,1296],"surroundings":[1187],"7B,":[1189,1312],"8A).":[1190,1313],"found":[1194],"coexisting":[1195],"Davidius":[1200],"sp.,":[1201],"Chlorogomphus":[1202],"spp.,":[1203,1205],"Heliogomphus":[1204],"Megalestes":[1206],"australis":[1207],"Karube,":[1208],"2014,":[1209],"Euphaea":[1210],"saola":[1211],"Hayashi":[1214],"Phan,":[1216],"Kompier,":[1217],"Karube":[1218],"Hayashi,":[1220],"2018,":[1221],"Cryptophaea":[1222],"vietnamensis":[1223],"(van":[1224],"Tol":[1225],"&Rozendaal,":[1226],"1995),":[1227],"Coeliccia":[1228],"scutellum":[1229],"Laidlaw,":[1230],"1932,":[1231],"Copera":[1233],"marginipes":[1234],"(Rambur,":[1235],"1842)":[1236],"(Phan":[1237],"2023,":[1240],"2024;":[1241],"2024).":[1245],"Some":[1246],"specimens":[1247],"had":[1248],"1\u20132":[1249],"chironomid":[1250],"attached":[1252],"space":[1255],"between":[1256,1261],"groove":[1260],"phoretic":[1268],"association":[1269],"7D\u2013E).":[1271],"exhibit":[1274],"specialized":[1276],"ambush":[1277],"predation":[1278],"strategy":[1279],"characterized":[1280,1319],"inhabiting":[1282],"muddy":[1283],"sandy":[1285],"substrates":[1286],"while":[1287,1333],"remaining":[1288],"partially":[1289],"buried":[1290],"motionless,":[1292],"blending":[1293],"surroundings.":[1297],"This":[1298],"behavior":[1299],"reduces":[1300],"detection":[1301],"predators":[1304],"prey,":[1306],"including":[1307],"instances":[1308],"cannibalism":[1310],"Additionally,":[1314],"they":[1315],"displayed":[1316,1338],"agonistic":[1317],"behaviour,":[1318],"tendency":[1322],"hold":[1324],"end":[1327],"abdomen":[1330],"upturned,":[1332],"appendage":[1336],"was":[1337],"upwards":[1341],"8B).":[1343]},"counts_by_year":[],"updated_date":"2026-10-14T16:39:01.820549","created_date":"2025-10-10T00:00:00"}